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John Morse

Professor Emeritus of Entomology
Director Emeritus of the Clemson University Arthropod Collection
Plant and Environmental Sciences Department

Office: 310 Long Hall
Phone: 864-656-5049
Fax: 8646563443

Personal Website:

 Educational Background

Ph.D. Entomology
University of Georgia 1974

M.S. Entomology
Clemson University 1970

B.S. Biology
Davidson College 1968

 Courses Taught

Aquatic Insects (1975-present)
Taxonomy of Immature Insects (1982-present)
Applied Systematics (1976-2014)
Creative Inquiry--Insects of the Clemson Experimental Forest (2001-present)
Biocomplexity (2002-present)


Besides courses in insect systematics, insect larvae, and a variety of other topics, I have taught courses related to aquatic insects at Clemson, at Highlands Biological Station in North Carolina, in several other states, and in 9 countries so far in East Asia for more than 42 years. My research specialty is the identification, biology, and historical development of caddisflies, which I have studied in many streams of the world since 1967. My students and I also investigate the identification, biology, and distribution of other aquatic insects, of stream ecology and conservation, and of the use of insect communities to monitor water pollution.

 Research Interests

World Trichoptera systematics and historical biogeography; Aquatic insect faunistics and biology; Use of insects for water quality biomonitoring; Insect species diversity and endangered species


136. Morse, J.C. 2009. Chapter 8: Biodiversity of aquatic insects. Pp. 165-184 in R.G. Foottit and P.H. Adler, editors, Insect Biodiversity: Science and Society. Wiley-Blackwell, Oxford, United Kingdom. 632 pp.

137. Morse, J.C. 2009. Macroinvertebrates as sentinels in freshwater biomonitoring. Pp. 171-173 in Priroda bez granits: Materiali III Mejdunarodnogo ekologicheskogo Foruma, Valdivostok, 12-13 Noyabrya 2008. Administratsiya Primorskogo Kraya, izd-vo Dalnyevost. un-ta, 2009. 477 pp.

138. Morse, J.C. 2009. Trichoptera (caddisflies). Pages 1015-1020 in V.H. Resh and R.T. Cardé (editors), Encyclopedia of Insects, 2nd edition. Academic Press, New York.

139. Hodges, A.C., & J.C. Morse. 2009. Southern Plant Diagnostic Network Invasive Arthropod Workshop, May 7–9, 2007. 13pp. Journal of Insect Science 9:61, available online:

140. Sun, C.-H., L.-F. Yang, & J.C. Morse. 2009. A new record genus and two new species of Arctopsychinae (Trichoptera: Hydropsychidae) from China. Acta Zootaxonomica Sinica 34(4): 912-916.

141. Chuluunbat, S., J.C. Morse, J.A.L. Lessard, M.E. Benbow, M.D. Wesener, & J. Hudson. 2010. Evolution of terrestrial habitat in Manophylax species (Trichoptera: Apataniidae), with a new species from Alaska. Journal of the North American Benthological Society 29(2): 413-430. Can be downloaded from TigerPrints:

142. Zhong, H., L.-F. Yang, & J.C. Morse. 2010. Four new species and two new records of Polyplectropus from China (Trichoptera: Polycentropodidae). Zootaxa 2428: 37-46. Open Access in Zootaxa: . Can be downloaded also from TigerPrints:

143. Geraci, C.J., X. Zhou, J.C. Morse, & K.M. Kjer. 2010. Defining the genus Hydropsyche (Trichoptera: Hydropsychidae) based on DNA and morphological evidence. Journal of the North American Benthological Society 29(3): 918-933. Can be downloaded from TigerPrints:

144. Zhou, L., L.-F. Yang, & J.C. Morse. 2010. Six new species and 1 new species record of Orthotrichia (Trichoptera: Hydroptilidae) from China. Zootaxa 2560: 29-41. Open Access in Zootaxa: . Can be downloaded also from TigerPrints:

145. Chuluunbat, S., Chuluunbaatar, G., & J.C. Morse. 2010. A redescription of Apatania mongolica Martynov, 1914 (Trichoptera: Apataniidae), based on materials from southern Mongolia. Mongolian Journal of Biological Sciences 8(1): 3-8.

146. Pfrender, M.E., C.P. Hawkins, M Bagley, G.W. Courtney, B.R. Creutzburg, J.H. Epler, S. Fend, L.C. Ferrington, Jr., P.L. Hartzell, S. Jackson, D.P. Larsen, C.A. Lévesque, J.C. Morse, M.J. Petersen, D. Ruiter, D. Schindel, & M Whiting. 2010. Assessing macroinvertebrate biodiversity in freshwater ecosystems: Advances and challenges in DNA-based approaches. The Quarterly Review of Biology 85(3): 319-340.

147. Oláh, J., & J.C. Morse. 2010. A synonym of Potamyia chekiangensis (Schmid 1965) (= P. chinensis Martynov, 1930 = P. martynovi Oláh & Morse, 2008). Braueria 37: 4.

148. Yang, L.-F., & J.C. Morse. 2010. Glossosoma (Lipoglossa) sinuatum Yang and Morse, nomen novum. Braueria 37: 4.

149. Majecka, K., J. Majecki, & J. Morse (Eds.). 2011. Proceedings of the 13th International Symposium on Trichoptera. Zoosymposia 5. 512 pp.

150. Majecka, K., J. Majecki, & J. Morse. 2011. PREFACE: Proceedings of the 13th International Symposium on Trichoptera. Pp. 7-10 in K. Majecka, J. Majecki, & J. Morse (Eds.), Proceedings of the 13th International Symposium on Trichoptera, Zoosymposia 5. 512 pp. Open Access in Zoosymposia: . Can be downloaded also from TigerPrints:

151. Morse, J.C. 2011. The Trichoptera World Checklist. Pp. 372-380 in K. Majecka, J. Majecki, & J. Morse (Eds.), Proceedings of the 13th International Symposium on Trichoptera, Zoosymposia 5. 512 pp. Open Access in Zoosymposia: . Can be downloaded also from TigerPrints:

152. Sun, C-h., L-f. Yang, & J.C. Morse. 2011. Three new species of Cheumatopsyche ( Trichoptera: Hydropsychidae) from Guangxi China. Journal of Nanjing Agricultural University 34(1): 57-61.

153. Holzenthal, R.W., J.C. Morse, & K.M. Kjer. 2011. Order Trichoptera Kirby, 1813. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 210-211. Open Access in Zootaxa: .

154. Morse, J.C., H. Zhong, & L.-f. Yang. 2012. New species of Plectrocnemia and Nyctiophylax (Trichoptera, Polycentropodidae) from China. ZooKeys 169: 39-59. Open Access in ZooKeys: .

155. Burington, Z.L., J.C. Morse, & J.V. McArthur. 2012. Distribution and variation of Oecetis parva (Trichoptera: Leptoceridae). Entomological News 122(1): 100-106.

156. Zhong, H., L.-f. Yang, & J.C. Morse. 2012. The genus Plectrocnemia Stephens in China (Trichoptera, Polycentropodidae. Zootaxa 3489: 1-24. Open Access in Zootaxa: . Can be downloaded also from TigerPrints:

157. Harvey, L.E., C.J. Geraci, J.L. Robinson, J.C. Morse, K.M. Kjer, & X. Zhou. 2012. Diversity of mitochondrial and larval morphological characters in the genus Diplectrona (Trichoptera: Hydropsychidae) in the eastern United States. Terrestrial Arthropod Reviews 5: 191-211.

158. Harrison, A.B. and J.C. Morse. 2012. The macroinvertebrate fauna of the Mississippi River. Transactions of the American Entomological Society 138: 35-52.

159. Stribling, J., B. Sweeney, J. Morse, G. Corkum, G. Lester, S. Miller, R. Mitchell, B. Poulton, S. Strachan, & M Wetzel. 2013. “Taxonomic certification versus the scientific method”: a rebuttal of Rogers (2012). Zootaxa 3359: 65–68.

160. Thawarorit, K., N. Sangpradub, & J.C. Morse. 2013. Five new species of the genus Cheumatopsyche (Trichoptera: Hydropsychidae) from the Phetchabun Mountains, Thailand. Zootaxa 3613 (5): 445–454. Open Access in Zootaxa: .

161. Chapin, J.W. & J.C. Morse. 2013. A new species of Micrasema (Trichoptera: Brachycentridae) from the Gulf Coastal Plain, U.S.A. Entomological News 123(2): 131-138.

162. Zhong, H., L.-F. Yang, & J.C. Morse. 2013. Pahamunaya Schmid, a new genus record from China, with description of a new species (Trichoptera: Polycentropodidae). Acta Zootaxonomica Sinica 38(2): 307-310.

163. Stark, B.P. & J.C. Morse. 2013. A new species of Neonemura (Plecoptera: Notonemouridae), and records of stoneflies from Aisén Province, Region XI, Chile. Illiesia, 9(10): 110-115.
Available online:

164. Morse, J.C., G.A. Nair, & S.S. Kumaran. 2013. New species of Ecnomidae (Trichoptera) from the southern Western Ghats, Kerala, India. In: K. Tojo, K. Tanida, & T. Nozaki (Eds.), Biology of Inland Waters, Supplement 2 (Proceedings of the 1st Symposium of the Benthological Society of Asia): 93-100.

165. Zhou, L., L.-f. Yang, & J.C. Morse. 2013. New species of Stactobia McLachlan (Trichoptera: Hydroptilidae) from China. Journal of the Kansas Entomological Society 86(3): 277-286. Download available from BioONE:

166. Ito, T., R.W. Wisseman, J.C. Morse, M.H. Colbo, & J.S. Weaver, III. 2014. The genus Palaeagapetus Ulmer (Trichoptera, Hydroptilidae, Ptilocolepinae) in North America. Zootaxa 3794 (2): 201–221. Open Access in Zootaxa: . Can be downloaded also from TigerPrints:

167. Mobley, C., C. Lee, J.C. Morse, J. Allen, & C. Murphy. 2014. Learning about sustainability: An interdisciplinary graduate seminar in biocomplexity. International Journal of Sustainability in Higher Education 15(1): 16-33.

168. Zhong, H., L.-f. Yang, & J.C. Morse 2014. The genus Nyctiophylax in China (Trichoptera, Polycentropodidae). Zootaxa 3846 (2): 273-284. Open Access in Zootaxa: . Can be downloaded also from TigerPrints:

169. Nair, G.A., J.C. Morse, & S A. Marshall. 2015. Aquatic insects and their societal benefits and risks. Keraljyothi 52(1): 7-11. [in Hindi]
Also, Journal of Entomology and Zoology Studies 3 (3): 171-177. [in English]

170. Zhou, L., L-f. Yang, & J.C. Morse. 2016. New species of microcaddisflies from China (Trichoptera: Hydroptilidae). Zootaxa 4097(2): 203-219.

171. Parey, S.H., J.C. Morse, & M.S. Pandher. 2016. Three new species of the genus Lepidostoma Rambur (Lepidostomatidae: Trichoptera) from India. Zootaxa 4136 (1): 181-187.

172. Zhou, X., P.B. Frandsen, R.W. Holzenthal, C.R. Beet, K.R. Bennett, R.J. Blahnik, N. Bonada, D. Cartwright, S. Chuluunat, G.V. Cocks, G.E. Collins, J. deWaard, J. Dean, O.S. Flint Jr., A. Hausmann, L. Hendrich, M. Hess, I.D. Hogg, B.C. Kondratieff, H Malicky, M.A. Milton, J. Morinière, J.C. Morse, F.N. Mwangi, S.U. Pauls, M R. Gonzalez, A. Rinne, J.L. Robinson, J Salokannel, M. Shackleton, B. Smith, A. Stamatakis, R. StClair, J.A. Thomas, Carmen Zamora-Muñoz, T. Ziesmann, & K.M. Kjer. 2016. The Trichoptera barcode initiative: A strategy for generating a species-level Tree of Life. Philosophical Transactions of the Royal Society B 371: 20160025.

173. Vshivkova, T.S., & J.C. Morse (Editors) 2016. Proceedings of the 14th International Symposium on Trichoptera. Zoosymposia 10: 1–507.
Vshivkova, T.S., & J.C. Morse. 2016. Preface: Proceedings of the 14th International Symposium on Trichoptera, Zoosymposia 10: 6–19.
Morse, J.C. 2016. Index to key words, taxa, geographical distribution, and authors. Zoosymposia 10: 502–507.

174. Morse, J.C. 2016. Keynote: The Trichoptera fauna of Asia. Zoosymposia 10: 20–28.

175. Chuluunbat, S., J.C. Morse, & S. Boldbaatar. 2016. Caddisflies of Mongolia: Distribution and diversity. Zoosymposia 10: 96–116.

176. Yang, L-f., C-h. Sun, & J.C. Morse. 2016. An amended checklist of the caddisflies of China (Insecta, Trichoptera). Zoosymposia 10: 451–479.

**High school student


Clemson University Arthropod Collection
Trichoptera World Checklist