Gregory Reighard

Professor of Pomology


Email: grghrd@clemson.edu
Office: 864-656-4962

Department:
School of Agricultural, Forest, and Environmental Sciences

Location:
161 Poole Agricultural Center, Box 340310


Education:
B.S.--Forest Science, Penn State University (1977)
M.S.--Biology, University of Michigan (1978)
Ph.D.--Forestry, Michigan State University (1984)

Research interest:
Development of Prunus rootstocks for resistance to nematodes, soil diseases and peach tree short life; Peach rootstock genetics and physiology emphasizing development of new rootstocks. Fruit tree culture and orchard floor management practices.

Publications:
Bell, R.L., T. van der Zwet, S. Castagnoli, T. Einhorn, J.D. Turneer, R. Spotts, G.A. Moulton, G.L. Reighard, and W. Shane. 2014. ‘Gem’ pear. HortScience (in press).
Frett, T. J., G. L. Reighard, W. R. Okie and K. Gasic. 2014. Mapping quantitative trait loci associated with blush in peach [Prunus persica (L.) Batsch]. Tree Genetics & Genomes
DOI: 10.1007/s11295-013-0692-y.
Zhebentyayeva, T. N., S. Fan, A. Chandra, D. G. Bielenberg, G. L. Reighard, W. R. Okie and A. G. Abbott. 2014. Dissection of chilling requirement and bloom date QTLs in peach using a whole genome sequencing of sibling trees from an F2 mapping population. Tree Genetics & Genomes 10:35-51.
Reighard, G.L. 2013. Peach, Plum and Apricot Rootstocks for the 21st Century. Aspects of Applied Biology 119:59-66.
Kenealy, L., G. Reighard, B. Rauh, W. Brideges, Jr.., and S. Gagnon. 2013. Spring temperatures influence growing degree day model harvest predictions for peach. Aspects of Applied Biology 119:197-200.
Yang, N, G. Reighard, D. Ritchie, W. Okie, and K. Gasic. 2013. Mapping quantitative trait loci associated with resistance to bacterial spot (Xanthomonas arboricola pv. pruni) in peach. Tree Genetics and Genomes 9(2):573-586.
Autio, W.R., T.L. Robinson, B. Black, T. Bradshaw, J.A. Cline, R.M. Crassweller, C.G. Embree, E.E. Hoover, R.S. Johnson, G. Lang, M.L. Parker, G.L. Reighard, J.R. Schupp, M. Stasiak, M. Warmund, and D. Wolfe. 2011. Performance of ‘Fuji’ and ‘McIntosh’ Apple Trees after 10 Years as Affected by Several Dwarf Rootstocks in the 1999 NC-140 Apple Rootstock Trial. J. Amer. Pomol. Soc. 65(2):2-20.
Autio, W.R., T.L. Robinson, B. Black, T. Bradshaw, J.A. Cline, R.M. Crassweller, C.G. Embree, E.E. Hoover, R.S. Johnson, G. Lang, M.L. Parker, R.L. Perry, G.L. Reighard, M. Stasiak, M. Warmund, and D. Wolfe. 2011. Performance of ‘Fuji’ and ‘McIntosh’ Apple Trees after 10 Years as Affected by Several Semidwarf Rootstocks in the 1999 NC-140 Apple Rootstock Trial. J. Amer. Pomol. Soc. 65(2):21-38.
Reighard, G.L., T. Beckman, B. Belding, B. Black, P. Byers, J. Cline, W. Cowgill, R. Godin, R. S. Johnson, J. Kamas, M. Kaps, H. Larsen, T. Lindstrom, M. Newall, D. Ouellette, R. Pokharel, L. Stein, K. Taylor, C. Walsh, D. Ward, and M. Whiting. 2011. Six-Year Performance of 14 Rootstocks at 11 Sites in the 2001 NC-140 Peach Trial. J. Amer. Pomol. Soc. 65(1): 26-41.
Johnson, S., R. Andersen, W. Autio, T. Beckman, B. Black, P. Byers, J. Cline, C. Chavez-Gonzalez, W. Cowgill, R. Godin, G. Greene, M. Kaps, J. Kamas, H. Larsen, T. Lindstrom, D. Miller, M. Newall, D. Ophardt, D. Ouellette, R. Parra-Quezada, R. Pokharel, G. Reighard, T. Robinson, J. Schupp, L. Stein, K. Taylor, C. Walsh, D. Ward, M. Warmund, and M. Whiting. 2011. Performance of the 2002 NC-140 Cooperative Peach Rootstock Planting. J. Amer. Pomol. Soc. 65(1): 17-25.
Liang, H., T. Zhebentyayeva, B. Olukolu, D. Wilde, G.L. Reighard and A. Abbott. 2010. Comparison of gene order in the chromosome region containing a TERMINAL FLOWER 1 homolog in apricot and peach reveals microsynteny across angiosperms. Plant Science 179:390-398.
Robinson, T., D. Wolfe, J. Masabni, R. Andersen, A, Azarenko, J. Freer, G. Reighard, P. Hirst, R. Hayden, and B. McCluskey. 2010. Performance of Plum Rootstocks with ‘Stanley’, ‘Valor’, ‘Veeblue’, and ‘Santa Rosa’ as the Scions in the 1991 NC-140 Multi-State Plum Trial. J. Amer. Pomol. Soc. 64(3): 173-182.
Baugher, T.A., J. Schupp, K. Ellis, J. Remcheck, E. Winzeler, R.duncan, S. Johnson, K. Lewis, G. Reighard, G. Henderson, M. Norton, A. Dhaddey, and P. Heinemann. 2010. String blossom thinner designed for variable tree forms increase crop load management efficiency in trials in four United States peach-growing regions. HortTechnology 20(2):409-414.
Jimenez, S., G. L. Reighard and D. G. Bielenberg. 2010. Gene expression of DAM5 and DAM6 is suppressed by chilling temperaturesand inversely correlated with bud break rate. Plant Molecular Biology 73:157-167
Jiménez S, Z. Li, G.L. Reighard, and D.G. Bielenberg. 2010. Identification of genes associated with growth cessation and bud dormancy release using a dormancy-incapable tree mutant. BMC Plant Biology 10(25).
Black, B.L., D. Drost, T. Lindstrom, J. Reeve, J. Gunnell and G. L. Reighard. 2010. A comparison of root distribution patterns among Prunus rootstocks. J. American Pomological Society 64 (1): 52-62.
Fan, F., D. G. Bielenberg, T.N. Zhebentyayeva, G. L. Reighard, W.R. Okie, D. Holland and A. G. Abbott. 2009. Mapping quantitative trait loci associated with chilling requirement, heat requirement and bloom date in peach (Prunus persica). New Phytologist doi: 10.1111/j.1469-8137.2009.03119.x
Li, Z., G.L. Reighard, A.G. Abbott and D. G. Bielenberg. 2009. Dormancy-associated MADS genes from the EVG locus of peach [Prunus persica (L.) Batsch] have distinct seasonal and photoperiodic expression patterns. J. Experimental Botany 60 (12): 3521-3530.
Olukolu, B. A., T. Trainin, S. Fan, C. Kole, D. G. Bielenberg, G. L. Reighard, A. G. Abbott, and D. Holland. 2009. Genetic linkage mapping for molecular dissection of chilling requirement and budbreak in apricot (Prunus armeniaca L.). Genome 52: 819-828.
Jimenez, S., A. L. Lawton-Rauh, G. L. Reighard, A. G. Abbott, and D. G. Bielenberg. 2009. Phylogenetic analysis and molecular evolution of the dormancy associated MADS-box genes from peach. BMC Plant Biology 2009, 9:81.
Okie, W.R., G.L. Reighard, and A.P. Nyczepir. 2009. Importance of Scion Cultivar in Peach Tree Short Life. J. American Pomological Society 63(2):58-63.
Bielenberg D.G., Y. Wang, Z. Li, T. Zhebentyayeva, S. Fan, G.L. Reighard, R. Scorza, and A.G. Abbott. 2008. Sequencing and annotation of the evergrowing locus in peach [Prunus persica (L.) Batsch] reveals a cluster of six MADS-box transcription factors as candidate genes for regulation of terminal bud formation. Tree Genomes & Genetics 4:495-507.
Liu, X., G.L. Reighard, G. Swire-Clark, and W.V. Baird 2008. Peach Rootstock Identification by DNA-fingerprinting with microsatellite (SSR) markers. Amer. Pom. Soc. 61: 162-166
Zhebentyayeva, T.N., G. Swire-Clark, L. Georgi, L. Garay, S, Jung, S. Forrest, A.V. Blenda, B. Blackmon, J. Mook, R. Horn, W. Howad, P. Arús, D. Main, J.P .Tomkins, B. Sosinski, W.V. Baird, G.L. Reighard and A.G. Abbott. 2008. A framework physical map for peach, a model Rosaceae species. Tree Genetics and Genomes DOI: 10.1007/s11295-008-0147-z.
Zhebentyayeva, T.N., G.L. Reighard, D. Lalli, V.M. Gorina, B.Krška, and A.G. Abbott. 2008. Origin of Resistance to Plum Pox Virus in Apricot: What New AFLP and Targeted SSR Data Analyses Tell. Tree Genetics and Genomes 4:403-417.
Njoroge, S. and G. L. Reighard. 2008. Thinning Time During Stage I and Fruit Spacing Influences Fruit Size of 'Contender' Peach. Scientia Horticulturae 115(4): 352-359.
Liu, X., G.L. Reighard, G.A. Swire-Clark and W.V. Baird. 2007. Peach rootstock identification by DNA-fingerprinting with microsatellite (SSR) markers. J. Amer. Pomol. Soc. 61(4): 162-166.
Blenda, A.V., I. Verde, L.L. Georgi, G.L. Reighard, S.D. Forrest, M. Muñoz-Torres, W.V. Baird, and A.G. Abbott. 2007. Construction of a genetic linkage map and identification of molecular markers in peach rootstocks for response to peach tree short life syndrome. Tree Genetics and Genomes 3:341-350.
Reighard, G., T. Beckman, R. Belding, B. Black, J. Cline, W. Cowgill, R. Godin, M. Kaps, T. Lindstrom. D. Ouellette, L. Stein, K. Taylor, C. Walsh, M. Whiting and T. Robinson. 2006. Performance of Prunus rootstocks in the 2001 NC-140 peach trial. Sodininkyste Ir Darzininkyste 25(3): 327-333.
Blenda, A.V., W.P. Wechter, G.L. Reighard, W.V. Baird and A.G. Abbott. 2006. Development and characterization of diagnostic AFLP markers in Prunus persica for its response to peach tree short life syndrome. J. Horticultural Sci. & Biotechnology 81(2):281-288.
Blenda, A.V., G.L. Reighard, W.V. Baird and A.G. Abbott. 2006. Simple sequence repeat markers for detecting sources of tolerance to PTSL Syndrome in Prunus persica rootstocks. Euphytica 147:287-295.
Marini, R.P., B.H. Barritt, G. R. Brown, J. Cline, W. P. Cowgill, Jr., R. M. Crassweller, P. A. Domoto, D. C. Ferree, J. Garner, G. M. Greene, C. Hampson, P. Hirst, M. M. Kushad, J. Masabni, E. Mielke, R. Moran, C. A. Mullins, M. Parker, R. L. Perry, J. P. Prive, G. L. Reighard, T. Robinson, C. R. Rom, T. Roper, J. R. Schupp, E. Stover and R. Unrath. 2006. Performance of ‘Gala’ apple on four semi-dwarf rootstocks: A ten-year summary of the 1994 NC-140 semi-dwarf rootstock trial. J. Amer. Pomol. Soc. 60(2): 58-68.
Marini, R.P., J. L. Anderson, W. R. Autio, B.H. Barritt, J. Cline, W. P. Cowgill, Jr., R. M. Crassweller, R.M. Garner, A. Gaus, R. Godin, G.M. Greene, C. Hampson, P. Hirst, M. M. Kushad, J. Masabni, E. Mielke, R. Moran, C. A. Mullins, M. Parker, R. L. Perry, J. P. Prive, G. L. Reighard, T. Robinson, C. R. Rom, T. Roper, J. R. Schupp, E. Stover and R. Unrath. 2006. Performance of ‘Gala’ apple on 18 dwarfing rootstocks: A ten-year summary of the 1994 NC-140 rootstock trial. J. Amer. Pomol. Soc. 60(2): 69-83.
Lalli, D.A., V. Decroocq, A.V. Blenda, V. Schurdi-Levraud, L. Garay, O. Le Gall, V. Damsteegt, G.L. Reighard, and A.G. Abbott. 2005. Identification and mapping of resistance gene analogs (RGAs) in Prunus: a resistance map for Prunus. Theor. Appl. Genet. 111: 1504-1513.
Wang, Y., G.L. Reighard, D.R. Layne, A.G. Abbott, and H. Huang 2005. Inheritance of AFLP markers and their use for genetic diversity analysis in wild and domesticated pawpaw [Asimina triloba (L.) Dunal].. J. Amer. Soc. Hort. Sci. 130(4): 561-568
Horn, R., A.C. Lecouls, A. Callahan, A.Dandekar, L. Garay, P. McCord, W. Howad, H. Chan, I. Verde, K. Ramaswamy, D. Main, S. Jung, L. Georgi, S. Forrest, J. Mook, T. Zhebentyayeva, Y. Yu, H.R. Kim, C. Jesudurai, B. Sosinski, P. Arus, W.V. Baird, D. Parfitt, G. Reighard, R. Scorza, J. Tomkins, R. Wing, and A.G. Abbott. 2005. Candidate gene database and transcript map for peach, a model species for fruit trees. Theor. Appl. Genet. 110:1419-1428.
Autio, W. R., T. L. Robinson, B. H. Barritt, J. A. Cline, R. M. Crassweller, C. G. Embree, D. C. Ferree, M. E. Garcia, G. M. Greene, E. E. Hoover, R. S. Johnson, K. Kosola, J. Masabni, M. L. Parker, R. L. Perry, G. L. Reighard, S.D. Seeley, and M. Warmund. 2005. Performance of ‘Fuji’ and ‘McIntosh’ apple trees after 5 years as affected by several semidwarf rootstocks in the 1999 NC-140 apple rootstock trial. J. Amer. Pom. Soc. 59 (4): 192-201.
Autio, W. R., T. L. Robinson, B. H. Barritt, J. A. Cline, R. M. Crassweller, C. G. Embree, D. C. Ferree, M. E. Garcia, G. M. Greene, E. E. Hoover, R. S. Johnson, K. Kosola, J. Masabni, M. L. Parker, R. L. Perry, G. L. Reighard, S.D. Seeley, and M. Warmund. 2005. Performance of ‘Fuji’ and ‘McIntosh’ apple trees after 5 years as affected by several dwarf rootstocks in the 1999 NC-140 apple rootstock trial. J. Amer. Pom. Soc. 59 (4): 202-214.
Wang, Y., G. L. Reighard, D. R. Layne, A. G. Abbott, and H. Huang. 2005. Inheritance of AFLP markers and their use for genetic diversity analysis in wild and domesticated pawpaw [Asimina triloba (L.) Dunal]. J. Amer. Soc. Hort. Sci. 130(4):561-568.
Reighard, G., R. Andersen, J. Anderson, W. Autio, T. Beckman, T. Baker, R. Belding, G. Brown, P. Byers, W. Cowgill, D. Deyton, E. Durner, A. Erb, D. Ferree, A. Gaus, R. Godin, R. Hayden, P. Hirst, S. Kadir, M. Kaps, H. Larsen, T. Lindstrom, N. Miles, F. Morrison, S. Myers, D. Ouellette, C. Rom, W. Shane, B. Taylor, K. Taylor, C. Walsh, and M. Warmund. 2004. Growth and yield of Redhaven peach on 19 rootstocks at 20 North American locations. J. Amer. Pomol. Soc. 58(4):174-202.
Bielenberg, D.G., Y. Wang, S. Fan, G.L. Reighard, R. Scorza, and A.G. Abbott. 2004. A deletion affecting several gene candidates is present in the peach Evergrowing mutant. J. of Heredity 95(5):436-444.
Gibson, P. G., G. L. Reighard, G. L. Powell, and T. C. Jenkins. 2004. Graft transmissible agents affect membrane fatty acid saturation during dormancy release in peach. J. Amer. Soc. Hort. Sci. 129(5):649-652.

Professional society activity:
• Member: American Society for Horticultural Science, 1985-2014; Southern Region of the American Society for Horticultural Science, 1987-2014; International Society for Horticultural Science, 1989-2014; American Pomological Society, 1987-2014; National Peach Council, 1987-2014; International Dwarf Fruit Tree Association, 1987-2014
• Current Society Offices/Committees: American Society for Horticultural Science (ASHS)- Elected member of the Executive Committee for the Southern Region-ASHS, 2012-2015; International Society for Horticultural Science (ISHS)-Scientific Committee and Editorial Board Member for ISHS 8th International Peach Symposium, Elected Section Chair for ISHS Peach Working Group, 2009-2013 and 2013-2017; American Pomological Society (APS)-Advisory Board Committee member for the APS. 1997-1999, 2005-2007, 2009-2014; International Dwarf Fruit Tree Association-Academic advisor and proposal reviewer for the Research Funding Committee, 1999-2014.

Honors and awards:
National Peach Council Lifetime Achievement Award. (2013)
International Fruit Tree Association Research Award for Contributions to the Tree Fruit Industry (2012)
Clemson University Agric. Expt. Station’s Godley-Snell Award for Excellence in Agricultural Research. (2008)
Clemson University Board of Trustees Award for Faculty Excellence. (Years 2006, 2008, 2009, 2012)
American Society of Horticultural Science - Southern Region Julian C. Miller, Sr. Distinguished Researcher Award. (2008)
Clemson University Extension Service Team Award. For Peach Extension Programs. (2007)
American Pomological Society’s 2007 U.P. Hedrick Award to Ph.D. Advisee, Ms. Xiaoyu Liu. (2007)
American Society of Horticultural Science Outstanding Fruit Cultivar Award. (2007)
American Pomological Society Shepard Award for best journal paper. (Years 2005, 2010)
National Peach Council Carroll R. Miller Award. (2002)
Co-inventor of PVPA No. 1994 issued May 1, 2001 for Guardian® Brand ‘BY520-9’ peach rootstock.
Co-owner of trademark issued in 1997 for the name Guardian® Brand ‘BY520-9’.

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