Faculty Profiles

Josep Rutllant

Faculty

Office: Hendrix Student Center
Email: jrutlla@clemson.edu

Education and Professional Certifications

NATO Postdoctoral Fellowship
Center for Animal Transgenesis and Germ Cell Research
University of Pennsylvania School of Veterinary Medicine
New Bolton Center, PA

Ph.D. Degree, in Veterinary Medicine
School of Veterinary Medicine, Autonomous University of Barcelona 

L.V. (D.V.M.) Degree
School of Veterinary Medicine, Autonomous University of Barcelona

Areas of Expertise

Veterinary Anatomy and Embryology Education

Active Student Learning

Curriculum Innovation

Animal Reproduction

Gamete Biology and Preservation Methods

Publications

Weakley, M., Rutllant, J. Chapter 3: Developing, Implementing and Utilizing a Willed Body Program in a Veterinary Curriculum. In: AAVMC Use of Animals in Veterinary Education Handbook. Pp. 17-21. AAVMC. https://doi.org/10.35542/osf.io/5k894

Khamas, W., Rutllant, J. External Features of Chicken. In: Anatomy and Histology of the Domestic Chicken. Khamas and Rutllant (Eds.). Pp. 1-17. John Wiley & Sons.

Khamas, W., Rutllant, J., Noriega, K. Skeletal System. In: Anatomy and Histology of the Domestic Chicken. Khamas and Rutllant (Eds.). Pp. 19-33. John Wiley & Sons.

Rutllant, J., Khamas, W. Digestive System. In: Anatomy and Histology of the Domestic Chicken. Khamas and Rutllant (Eds.). Pp. 49-69. John Wiley & Sons.

Khamas, W., Rutllant, J. Urinary System. In: Anatomy and Histology of the Domestic Chicken. Khamas and Rutllant (Eds.). Pp. 83-92. John Wiley & Sons.

Rutllant, J., Khamas, W. Reproductive System. In: Anatomy and Histology of the Domestic Chicken. Khamas and Rutllant (Eds.). Pp. 93-108. John Wiley & Sons.

Khamas, W., Rutllant, J. Sense Organs. In: Anatomy and Histology of the Domestic Chicken. Khamas and Rutllant (Eds.). Pp. 125-137. John Wiley & Sons.

Khamas, W., Rutllant, J. The Lymphatic System. In: Anatomy and Histology of the Domestic Chicken. Khamas and Rutllant (Eds.). Pp. 153-170. John Wiley & Sons.

Khamas, W., Rutllant, J., Smodlaka, H.  Nervous System. In: Anatomy and Histology of the Domestic Chicken. Khamas and Rutllant (Eds.). Pp. 171-182. John Wiley & Sons.

Rutllant, J., Khamas, W. The Egg Anatomy. In: Anatomy and Histology of the Domestic Chicken. Khamas and Rutllant (Eds.). Pp. 209-216. John Wiley & Sons.

Rutllant, J., Khamas, W. Fertilization and Chick Embryo Development. In: Anatomy and Histology of the Domestic Chicken. Khamas and Rutllant (Eds.). Pp. 217-226. John Wiley & Sons.

Khamas W. Rutllant J. Atlas of the Anatomy and Histology of the Domestic Chicken. Wael Khamas and Josep Rutllant (Editors). Wiley (Publisher). Author and coauthor of 10 chapters of the book.

Khamas W. Rutllant J. Gamebird Anatomy. In: “Gamebird Medicine and Management”. Robert Porter and Teresa Morishita (Eds.)

Khamas W. Rutllant J. Anatomy and Physiology. In: “Backyard Poultry Medicine and Surgery: A Guide for Veterinary Practitioners”. Greenacre and Morishita (Eds.). Nov 2020 (2nd Edition). ISBN: 9781118335437

Khamas W. Rutllant J. and Greenacre C. Anatomy, Relevant Physiology and Physical Examination. In: “Backyard Poultry Medicine and Surgery: A Guide for Veterinary Practitioners”. Greenacre and Morishita (Eds.). Nov 2014. ISBN: 9781118335437

Morishita T. Karcher D. Rutllant J. Egg Diagnostics. In: “Backyard Poultry Medicine and Surgery: A Guide for Veterinary Practitioners”. Greenacre and Morishita (Eds.). Nov 2014. ISBN: 9781118335437

López-Gatius, F.; Rutllant, J.; López-Béjar, M.A. Management of Reproduction in Cattle and Buffaloes: The Bull and Artificial Insemination. CAB International. ISBN: 0851996248

Publications: Refereed Journals         

Wyatt S and Rutllant J. Anomalous branching of the aortic arch in a dog. Anat. Histol. Embryol. 51:347-354.

Betancourt, S., Irizarry, K., Falk, B., Rutllant, J., Khamas, W. Micromorphological study of the upper digestive tract of the Argentine tegu (Salvator merianae). Anat. Histol. Embryol. 51:259-268.

Irizarry K and  Rutllant J. Leveraging Comparative Genomics to Identify and Functionally Characterize Genes Associated with Sperm Phenotypes in Python bivittatus (Burmese Python). Genet Res Int. 2016: 7505268. PMCID: PMC4855019

Klein C, Troedsson M, Rutllant J. Region specific expression of aquaporin subtypes in equine testis, epididymis, and ductus deferens. Anat. Rec. 296(7):1115-26.

Klein C, Troedsson M, Rutllant J. Expression of Aquaporin Water Channels in Equine Endometrium is Differentially Regulated During the Oestrous Cycle and Early Pregnancy. Reprod. Dom. Anim. 48(4):529-37.

Klein C, Rutllant J, Troedsson M.H.T.  Expression Stability of Putative Reference Genes in Equine Endometrial, Testicular, and Conceptus Tissues. BMC Research Notes (BioMed Central) 4:120.

Rutllant, J, and, Meyers, S.A. Evidence for Membrane Raft formation during low temperature incubation of Macaque (Macaca mulatta) Spermatozoa. J. Androl. 30: 134-.

Rutllant, J; Alvarado, O; Fonda, E. Effects of trehalose on the osmotic stress limits of boar sperm. Biol. Reprod.   78:145-.

DeCarlo, C., Alderson, J., Bowen, E., Rutllant, J., Aldridge, B. Characterization of the Immunomodulatory Effect of a Commercial Extender on Semen-Induced Inflammatory Response in Equine Uterine Luminal Immune Cells. J. Reprod. Immunol. 71:158.

Rutllant, J.; López-Gatius, F; López-Béjar, M. The bovine vaginal fluid: a possible morphofunctional model for studying sperm transport in mammals. Reprod. Dom. Anim. 40:79-86.

Rutllant, J.; Pommer, A. C.; Meyers, S. A. Osmotic tolerance limits and properties of rhesus monkey (Macaca mulatta) spermatozoa. J. Androl. 24:534-541.

Pommer, A. C.; Rutllant, J.; Meyers, S. A. Phosphorylation of protein tyrosine residues in fresh and cryopreserved stallion spermatozoa under capacitating conditions. Biol. Reprod. 68:1208-1214.

Mayor, P.; López-Béjar, M.; Jori, F.; Rutllant, J.; López-Gatius, F; López-Plana, C. Anatomicohistological Characteristics of the Genital Tubular Organs of the Female Brush-tailed Porcupine (Atherurus africanus, Gray 1842) from Gabon. Anatomia, Histologia, Embryologia, 31:355-361.

Rutllant, J.; López-Béjar, M.; Santolaria, P.; Yániz, J.; López-Gatius, F. Rheological and ultrastructural properties of bovine vaginal fluid obtained at oestrus. J. Anat., 201:53-60.

Pommer, A. C.; Rutllant, J.; Meyers, S. A. The role of osmotic resistance on equine spermatozoal function. Theriogenology, 58:1373-1384.

Yániz, J.; López-Béjar, M. Santolaria, P.; Rutllant, J. ;López-Gatius, F. Intraperitoneal insemination in mammals: A review. Reprod. Dom. Anim., 37:75-80.

López-Gatius, F.; Santolaria, P.; Yániz, J.; Rutllant, J.; López-Béjar, M. Factors affecting pregnancy loss from gestation day 38 to 90 in lactating dairy cows from a single herd. Theriogenology, 57:1251-1261.

López-Gatius, F.; Santolaria, P.; Yániz, J.; Rutllant, J.; López-Béjar, M. Persistent ovarian follicles in dairy cows. A therapeutic approach. Theriogenology, 56:649-659.

Rutllant, J.; Meyers, S. A. Post-translational processing of PH-20 during epididymal sperm maturation in the horse.  Biol. Reprod. 65:1324-1331.

Rutllant, J.; López-Béjar, M.;Camón, J.; Martí, M.; López-Gatius, F. Confocal scanning laser microscopy of the bovine vaginal fluid at oestrus. Anatomia, Histologia, Embryologia, 30:159-162.

Ballester, J.; Fernández-Novell, J.M.; Rutllant, J.; García-Rocha, M.; Jesús Palomo, M.; Mogas, T.; Peña, A.; Rigau, T.; Guinovart, J.J.; Rodríguez-Gil, J.E. Evidence for a functional glycogen metabolism in mature mammalian spermatozoa. Molecular Reproduction and Development, 56(2):207-19.

Rutllant, J.; López-Gatius, F.; Camón, J.; López-Béjar, M; López-Plana, C. A structural study of the bovine vaginal fluid at estrus. Scanning, 21:204-211.

Villamediana, P.; Rutllant, J.; López-Béjar, M.; Vidal, F.; Paramio, MT. Changes in zona pellucida surface after in vivo and in vitro maturation of caprine oocytes. Reproduction in Domestic Animals, 34:417-421.

López‑Béjar, M. A.; López‑Gatius, F.; Camón, J.; Rutllant, J.; Valls, X.; Labèrnia, J.; Santolaria, P. Morphological features and effects on reproductive parameters of ovarian cysts of ovarian origin in superovulated rabbit does. Reproduction in Domestic Animals, 33:369-378.

LópezGatius, F.; Rutllant, J.; Labèrnia, J.; Ibarz, A.; LópezBéjar, M.A.; Santolaria, P. The relationship of rheological behavior of the vaginal fluid at the time of insemination in dairy cows.  Theriogenology, 48:865-871.

López‑Béjar, M. A.; López‑Gatius, F.; Camón, J.; Rutllant, J.; Labèrnia, J.; Santolaria, P.  Rapid freezing of rabbit embryos has a negative effect on embryo morphology.  Reproduction in Domestic Animals, 32:237-241.

Rutllant, J.: LópezGatius, F.; Camón, J.; LópezPlana, C.; LópezBéjar, M. A.  A scanning electron microscope study of the structural component of the bovine vaginal fluid at oestrus.  Journal of Veterinary Medicine, Series A, 44:237-241.

López‑Béjar, M. A.; López‑Gatius, F.; Rutllant, J.; Camón, J.; Labèrnia, J.; Santolaria, P. Development in vivo of rabbit morulae after freezing using a two‑step cooling method. Reproduction in Domestic Animals, 31:661-664.

 LópezGatius, F.; Labèrnia, J.; Santolaria, P.; LópezBéjar, M.A.; Rutllant, J.  Effect of reproductive disorders previous to conception on pregnancy attrition in dairy cows.  Theriogenology, 46:643648.

LópezGatius, F.; Rutllant, J.; Labèrnia, J.; Ibarz, A.; LópezBéjar, M.A.; Santolaria, P. Rheological behavior of the vaginal fluid of dairy cows at estrus.  Theriogenology, 46:5763.

Labèrnia, J.; LópezGatius, F.; Santolaria, P.; LópezBéjar, M.A.; Rutllant, J.  Influence of management factors on pregnancy attrition in Dairy cattle.  Theriogenology, 45:12471253.

Camón, J.; LópezBéjar, M. A.; Verdú, J.; Rutllant, J.; Sabaté, D.; Degollada, E.; López-Plana, C.  Persistent truncus arteriosus in a diprosopic newborn calf.  Journal of Veterinary Medicine, Series A 42:4149.

Select Honors and Awards

Zoetis-Distinguished Teaching Award
Western Carolina University’s College of Veterinary Medicine
2025

University Distinguished Teacher Award
Western University of Health Sciences
2024

Outstanding Faculty Award Class of 2025
Western Carolina University Student Government Association
2022

Zoetis-Distinguished Teaching Award
Western Carolina University’s College of Veterinary Medicine
2015

Pfizer Award for Research Excellence
Western Carolina University’s College of Veterinary Medicine
2010

Norden-Pfizer Distinguished Teaching Award
Western Carolina University’s College of Veterinary Medicine
2006

Teacher of the year, Class of 2006
School of Veterinary Medicine, University of California Davis
2002–2003