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Clemson University
college of agriculture, forestry and life sciences clemson university

G. Geoff Wang

Forestry and Environmental Conservation Department, Experiment Station

Office: 226 Lehotsky Hall
Phone: 864-656-4864
Fax: 864-656-3304

Vita: Download CV


Educational Background

Ph.D. Forest Ecology
Univ. of British Columbia

Courses Taught

Silviculture Laboratory
Fire Ecology and Management
Fire Ecology and Management Laboratory
Creative Inquiry (Undergraduate Research)
Selected Topics in Forest Resources
Master Thesis Research
Doctoral Dissertation Research


Dr. Wang directs the Silviculture and Ecology Lab (SEL) at Clemson University. He has been either PI/ Project Leader or Co-PI for ~65 externally funded projects. Currently, Dr. Wang has four ongoing research projects, including an active program of national and international research collaborations. He has published ~176 peer-reviewed journal articles and over ~75 non-refereed publications. He has also delivered ~280 conference presentations, including ~80 invited seminars. His research has made a great impact in the field, as indicated by ~5200 citations and an h-index of 39 (Google Scholar).

Dr. Wang has taught multiple undergraduate graduate courses and mentored 21 post-doctoral scientists, 15 doctoral students, 22 master students, and seven honors students. Students under his mentorship have won many prestigious scholarships and awards. Many Ph.D. students and post-docs trained in his lab currently hold tenure-track faculty positions or equivalent.

Dr. Wang currently serves as an Associates Editor for New Forests, Canadian Journal of Forest Research, and Journal of Forestry Research; as a member of the editorial board for Agricultural and Rural Studies, Journal of Plant Ecology, and Journal of Forests and Environment. Dr. Wang has been frequently consulted nationally and internationally by scientific journals, grant agencies, government agencies, professional forestry associations, and industries. Internally, he has also served and chaired committees at departmental, college and university levels.

Research Interests

Dr. Wang specializes in Silviculture and Forest Ecology. He conducts both applied and foundational ecological research in forest ecosystems, mostly in temperate forests. His current research focuses on climate-smart forestry and sustainable forest management, especially on how natural and anthropogenic disturbances, including fire, weather extremes, pest/disease, biological invasion, and human activities (including management), affect forest ecosystems and their implication for forest resilience and sustainability.


Refereed Research Articles since 2019 (n = 56; updated October, 2023)
(*denote the corresponding author; +denote the leading author who completed the whole or a portion of the work in my lab)

Yang, K., Q. Zhang, J.J. Zhu, Q. Wang, T. Gao, and G.G Wang. 2023. Mycorrhizal associations drive the tradeoff between plant forest biomass and soil carbon accumulations in forest ecosystems under N deposition and warming. Nature Climate Change (Accepted)

Pile Knapp, L.S., D.R. Coyle, D.C., Dey, J.S. Fraser, T. Hutchinson, M.A. Jenkins, C.C. Kern, B.O. Knapp, D. Maddox, C. Pinchot, and G.G. Wang. 2023. Invasive plant management in eastern North American Forests: A systematic review. Forest Ecology and Management, 550, 121517.

Han, W., L. Chen, J. Liu, G.G. Wang, D. Liu, and G. Liu. 2023. Effect of Gap Size and Elevation on the Regeneration and Coexistence of Abies, Betula, and Acer Tree Species in a Subalpine Coniferous Forest. Forests, 14(10), 2099.

Tong, R., C. Ma, C. Lou, W. Yuan, N. Zhu, G.G. Wang, and T.G. Wu. 2023. Leaf nitrogen and phosphorus stoichiometry of the halophytes across China. Frontiers in Plant Science, 14, 1276699.

Lu, D.L., J.J. Zhu, G. Zhang, S. Fang, Y. Sun, C. Zhu, J. Zhang, and G.G. Wang. 2023. A forest gap is not forever: Towards an objective standard to determine when a gap is considered closed in temperate forests. Agricultural and Forest Meteorology 340,

Tang+, Y., X. Liu, J. Lian, X. Cheng, G.G. Wang, and J. Zhang. 2023. Soil Depth Can Modify the Contribution of Root System Architecture to the Root Decomposition Rate. Forests, 14(6), 1092.

Wang, J-P., R-Z. Huang*, J-M. Zhao, J-H. Yuan, G-M. Huang, R-X. Xie, L. Hu, Y-F. Xie, and G.G. Wang*. 2023. Effects of cultivars and habitats on rhizospheric arbuscular mycorrhizal fungal communities associated with Gannan navel oranges. Archives of Agronomy and Soil Science (GAGS),

Lu, D.L., J.J. Zhu, G. Zhang, Y. Sun, L. Hu, and G.G. Wang. 2023. Disentangling regeneration by vertical stratification: A 17-year gap-filling process in a temperate secondary forest. Forest Ecology and Management, 539: 120994,

Chi, Y., G.G. Wang, M. Zhu, P. Jin, Y. Hu, P. Shu, Z. Wang, A. Fan, P. Qian, Y. Han, and S. Jin. 2023. Potentially suitable habitat prediction of Pinus massoniana Lamb. in China under climate change using Maxent model. Frontiers in Forests and Global Change, 6.

Qi, K., J.J. Zhu, X. Zheng, G.G. Wang, and M. Li. 2023. Impacts of the World’s Largest Afforestation Program (Three-North Afforestation Program) on desertification control in sandy land of China. Geoscience and Remote Sensing, 60(1),


Wang, J., Y. Liu, Y. Xu, W. Chen, Y. Han, G.G. Wang*, and S. Jin*. 2022. Sexual differences in gas exchange and chlorophyll fluorescence of Torreya grandis under drought stress. Trees 36, 283–294.

Wang+, J., J. Yuan, Q. Ren, B. Zhang, J.-C. Zhang, R. Huang, and G.G. Wang. 2022. Arbuscular mycorrhizal fungi enhanced salt tolerance of Gleditsia sinensis by modulating antioxidant activity, ion balance and P/N ratio. Plant Growth Regulator 97, 33–49.

Huang,Y-X., Z-J. Wu, Y-Y. Zong, W-Q. Li, F-S. Chen, and GG Wang, J-J Li, X-M Fang. 2022. Mixing with coniferous tree species alleviates rhizosphere soil phosphorus limitation of broad-leaved trees in subtropical plantations. Soil Biology and Biochemistry, 175, 108853,

Xie+, H.T., L.S. Pile Knapp, M. Yu, and G.G. Wang. 2022. Morphological and physiological response of Chinese tallow (Triadica sebifera) to an extreme cold spell in subtropical, coastal forests of China. Frontiers in Ecology and Evolution. 10: 945886.

Xie+, HT, L.S. Pile Knapp, M. Yu, and G.G. Wang. 2022. Solidago canadensis invasion destabilizes the understory plant community and soil properties of coastal shelterbelt forests of subtropical China. Plant and Soil. 484, 65–77

Zhang, H, M Sun, Y. Wen, R Tong, GG Wang, Q Wu, Y Li, and TG Wu. 2022. The effects of stand age on leaf N:P cannot be neglected: A global synthesis. Forest ecology and management, 518, (2022): 120294.

Tong R., Y. Wen, J. Wang, C. Lou, C. Ma, N. Zhu, G.G. Wang, and T.G. Wu. 2022. Root nutrient capture and leaf resorption efficiency modulated by different influential factors jointly alleviated P limitation in Quercus acutissima across the North–South Transect of Eastern China. Forestry Research 2:7

Yan, Q.L., C.Y. Zhu, J.J. Zhu, L.N. Song, G.G. Wang, and D.L. Lu. Estimating gap age using tree-ring width in combination with carbon isotope discrimination in a temperate forest, Northeast China. Annals of Forest Science, 79:25.

Song R., R. Tong, H. Zhang, G.G. Wang, T.G. Wu, and X. Yang. 2022. Effects of Long-Term Fertilization and Stand Age on Root Nutrient Acquisition and Leaf Nutrient Resorption of Metasequoia glyptostroboides. Frontiers in Plant Science. 13

Li, W-Q., Z-J. Wu, Y-Y. Zong, G.G. Wang, F-S. Chen, Y-Q. Liu, J-J. Li, X-M. Fang. 2022. Tree species mixing enhances rhizosphere soil organic carbon mineralization of conifers in subtropical plantations, Forest Ecology and Management, 516, 120238,

Yang, K., J.J. Zhu, W. Zhang, Q. Zhang; D.L. Lu, Y. Zhang, X. Zheng, S. Xu, and G.G. Wang. 2022. Litter decomposition and nutrient release from monospecific and mixed litters: comparisons of litter quality, fauna and decomposition sites effects. Journal of Ecology

Wang+, J., G.G. Wang, R. Li, Y. Sun, L. Yu, J.J. Zhu, and Q. Yan. 2022. Cascading effect of source limitation on the granivore-mediated seed dispersal of Korean pine (Pinus koraiensis) in secondary forest ecosystems. Ecological Processes 11 (1),

Crouch, C.D., B.O. Knapp, S.A. Cohen, J.S. Glitzenstein, J.L. Walker, and G.G. Wang. 2022. Longleaf pine restoration on hydric sites: Understorey plant community responses to site preparation through 15 years. Applied Vegetation Science 25 (1), e12637


Wen, Y., R. Tong, H. Zhang, K. Feng, R. Song, G.G. Wang*, and T.G. Wu*. 2021. N addition decreased stand structure diversity in young but increased in middle-aged Metasequoia glyptostroboides plantations. Global Ecology and Conservation 30,

Wang+, K., G.G. Wang, L. Song, R. Zhang, T. Yan, and Y. Li. 2021. Linkages between nutrient resorption and ecological stoichiometry and homeostasis along a chronosequence of Mongolian pine plantations. Frontiers in Plant Science 13,

Han, Y., G.G. Wang, T.G. Wu, W. Chen, Y. Ji, and S. Jin. 2021. Fertilization failed to make positive effects on Torreya grandis in severe N-deposition subtropics. Sustainability 13, 9736.

Zhang, T., Q.L. Yan*, G.G. Wang, and J.J. Zhu. 2021. The effects of stump size and within-gap position on sprout non-structural carbohydrates concentrations and regeneration in forest gaps vary among species with different shade tolerances. Ecological Processes 10(25),

Wu+, C., M.D. Ulyshene, C. Shu, Z. Zhang, Y. Zhang, Y. Liu*, and G.G. Wang*. 2021. Stronger effects of termites than microbes on wood decomposition in a subtropical forest. Forest Ecology and Management 493, 119263,

Wang+, J., G.G. Wang*, T. Zhang, J. Yuan, L. Yu, J.J. Zhu, and Q.L. Yan*. 2021. Use of direct seeding and seedling planting to restore Korean pine (Pinus koraiensis Sieb. et Zucc.) in secondary forests of Northeast China. Forest Ecology and Management 493, 119243,

Lu+, D.L., J.J. Zhu, X. Wang, G. Hao, and G.G. Wang. 2021. A systematic evaluation of gap size and within-gap position effects on seedling regeneration in a temperate secondary forest, Northeast China. Forest Ecology and Management 490, 119140,

Xie*+, H.T., Y. Tang, M.K. Yu, and G.G. Wang*. 2021. The effects of afforestation tree species mixing on soil organic carbon stock, nutrients accumulation, and understory vegetation diversity on reclaimed coastal lands in Eastern China. Global Ecology and Conservation 26 e01478.

Wang, J., S. Ma, G.G. Wang, L. Xu, Z. Fu, J. Song, and J. Zhang. 2021. Arbuscular mycorrhizal fungi communities associated with wild plants in a coastal ecosystem. Journal of Forestry Research 32: 683-695.

Zhu, J.J., C.Y. Zhu, D.L. Lu, G.G. Wang, X. Zheng, J.S. Cao, J.X. Zhang. 2021. Regeneration and succession: A 50-year gap dynamic in temperate secondary forests, Northeast China. Forest Ecology and Management 484: 118943

Fang, X.-M. G.G. Wang, Z.-J. Xu, Y.-Y. Zong, X.-L. Zhang, J.-J. Li, H. Wang, F.-S. Chen. 2021. Litter addition and understory removal influenced soil organic carbon quality and mineral nitrogen supply in a subtropical plantation forest. Plant and Soil 460, 527–540.


Lu, D.L. J.J. Zhu, D. Wu, Q. Chen, Y. Yu, J. Wang, C. Zhu, H. Liu, T. Gao, G.G. Wang. 2020. Detecting dynamics and variations of crown asymmetry induced by natural gaps in a temperate secondary forest using terrestrial laser scanning. Forest Ecology and Management 473(118289)

Lu+ D.L., L.S. Pile, D.P. Yu, J.J. Zhu, D.C. Bragg, G.G. Wang*. 2020. Differential responses of tree species to a severe ice storm and their implications to forest composition in the southeast United States. Forest Ecology and Management 468(118177)

Xie+, H.T., J.E. Fawcett, G.G. Wang*. 2020. Fuel dynamics and its implication to fire behavior in loblolly pine-dominated stands after southern pine beetle outbreak. Forest Ecology and Management 466(118130)

Wang+, J.P., L. Zhai, J. Ma, J. Zhang*, G.G. Wang*, X. Liu, S. Zhangd, J. Songa, Y. Wu. 2020. Comparative physiological mechanisms of arbuscular mycorrhizal fungi in mitigating salt-induced adverse effects on leaves and roots of Zelkova serrata. Mycorrhiza. 30: 341-355.

Zhang, P. Y. Wen, L. Wang, H. Zhang, G.G. Wang, T.-G. Wu. 2020. Leaf structural carbohydrate decreased for Pinus thunbergii along a coast-inland gradient. Forests, 11, 449.

Wang+, J., S. Ma, G.G. Wang, L. Xu, Z. Fu, J. Song, J.C. Zhang. 2020. Arbuscular mycorrhizal fungi communities associated with wild plants in a coastal ecosystem. J. For. Res.

Xiao+, S.S., G.G. Wang, Tang C., Fang, H., Duan J., Yu X.F*. 2020. Effects of One-Year Simulated Nitrogen and Acid Deposition on Soil Respiration in a Subtropical Plantation in China. Forests,11(2): 235.

Crouch, C.D., B.O. Knapp, S.A. Cohen, M.C. Stambaugh, J.L.Walker, and G.G. Wang. 2020. Site preparation for longleaf pine restoration on hydric sites: Stand development through 15 years after planting. Forest Ecology and Management 461

Wu+, C., Z. Zhang, C. Shua, Q. Mo, H. Wang, F. Kong, Y. Zhang, G.G. Wang*, and Y. Liu*. 2020. The response of coarse woody debris decomposition and microbial community to nutrient additions in a subtropical forest. Forest Ecology and Management 460


Jin+, S.H., B.M. Moule, D. Yu, and G.G. Wang*. 2019. Fire survival of longleaf pine (Pinus palustris) grass stage seedlings: the role of seedling size, root collar position, and resprouting. Forests 10, 1070.

Wang, J.N., J.Y. Wang, L Wang, H. Zhang, Z. Guo, G.G. Wang, W.K. Smith, T.G. Wu. 2019. Does stoichiometric homeostasis differ among tree organs and with tree age? Forest Ecology and Management 453:117637.

Zhang, X., B. Zhang, K. Feeley, G.G. Wang, J. Zhang, and L. Zhai. Ecological contingency in species shifts: Downslope shifts of wood species under warming climate and land-use change. Environmental Research Letters 14: 114033.

Lu. D., G. Zhang, J.J. Zhu, G.G. Wang, C. Zhu, Q.L. Yan, J. Zhang. 2019. Early natural regeneration patterns of woody species within gaps in a temperate secondary forest. European Journal of Forest Research

Wang+, F., X.-M. Fang, G.G. Wang*, R. Mao, X.-F. Lin, H. Wang, F.-S. Chen*. 2019. Effects of nutrient addition on foliar phosphorus fractions and their resorption in different-aged leaves of Chinese fir in subtropical China. Plant and Soil

Wang+, J.P. Z. Zhong, L. Zhu, Y. Yuan, L. Xu, G.G. Wang, L. Zhai, L. Yang, and J. Zhang. 2019. Arbuscular Mycorrhizal fungi effectively enhance the growth of Gleditsia sinensis Lam. seedlings under greenhouse conditions. Forests 10, 567; doi:10.3390/f10070567

Ye, X.M., Y. Zhang, F.S. Chen, G.G. Wang, X.M Fang, X.F. Lin, S.Z. Wan, and P. He. 2019. The effects of simulated deposited nitrogen on nutrient dynamics in decomposing litters across a wide quality spectrum using a 15-N tracing technique. Plant and Soil

Zhu, C., J.J. Zhu, G.G. Wang, X. Zheng, D. Lu, and T. 2019. Dynamics of gaps and large openings in a secondary forest of Northeast China over 50 years. Annals of Forest Science

Wang, J.-P., G.G. Wang, B. Zhang, Z. Yuan, Z. Fu, Y. Yuan, L. Zhu, S. Ma, and J. Zhang. 2019. Arbuscular mycorrhizal fungi associated with tree species in a planted forest of Eastern China. Forests 10(5):424 DOI: 10.3390/f10050424

Pile+, L.S., L. Vickers, M. Stambaugh, C. Norman, and G.G. Wang*. 2019. The Tortoise and the Hare: A Race between Native Tree Species and the Invasive Chinese Tallow. Forest Ecology and Management 445: 110-121.

Wu+, C., Z. Zhang, H. Wang, G. Huang, C. Shu, F. Kong, Y. Zhang, G.G. Wang*, and Y. Liu*. 2019. Home-field advantage of CWD decomposition in subtropical forests varied by field sites. Forest Ecology and Management 444: 127-137.

Chen* F.-S., G.G. Wang*, X.-M. Fang, S.-Z. Wan, Y. Zhang, and C. Liang. 2019. Nitrogen deposition effect on forest litter decomposition is interactively regulated by endogenous litter quality and exogenous resource supply. Plant and Soil 437: 413-426.

Wang, F., F.-S. Chen, G.G. Wang, R. Mao, X. Fang, H. Wang, and W. Bu. 2019. Effects of experimental nitrogen addition on nutrients and nonstructural carbohydrates of dominant understory plants in a Chinese fir plantation. Forests 10, 155; doi:10.3390/f10020155

Wu+, C., H. Wang, Q. Mo, Z. Zhang, G. Huang, F. Kong, Y. Liu*, G.G. Wang*. 2019. Effects of elevated UV-B radiation and N deposition on the decomposition of coarse woody debris. The Science of Total Environment 663:170-176.


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