G. Geoff Wang

Forestry and Environmental Conservation Department, Experiment Station

Office: 226 Lehotsky Hall
Phone: 864-656-4864
Fax: 864-656-3304

Email: gwang@clemson.edu

 Educational Background

Ph.D. Forest Ecology
Univ. of British Columbia

 Courses Taught

Silviculture Laboratory
Fire Ecology and Management
Fire Ecology and Management Laboratory
Creative Inquiry (Undergraduate Research)
Selected Topics in Forest Resources
Master Thesis Research
Doctoral Dissertation Research


Dr. Wang directs the Silviculture and Ecology Lab (SEL) at Clemson University. He has been either PI/ Project Leader or Co-PI for ~60 externally funded projects. Currently, Dr. Wang has eight ongoing research projects, including an active program of national and international research collaborations. He has published ~150 peer-reviewed journal articles and over ~70 non-refereed publications. He has also delivered ~260 conference presentations, including ~70 invited seminars. His research has made a great impact in the field as indicated by ~3700 citations and an h-index of 33 (Google Scholar).

Dr. Wang has taught multiple undergraduate graduate courses and mentored 21 post-doctoral scientists, 15 doctoral students, 22 master students, and seven honors students. Students under his mentorship have won many prestigious scholarships and awards. Many Ph.D. students and post-docs trained in his lab currently hold tenure-track faculty positions or equivalent.

Dr. Wang currently serves as an Associates Editor for New Forests, Canadian Journal of Forest Research, and Journal of Forestry Research; as a member in the editorial board for Journal of Plant Ecology and Journal of Forests and Environment. Dr. Wang has been frequently consulted nationally and internationally by scientific journals, grant agencies, government agencies, professional forestry associations, and industries. Internally, he has also served and chaired committees at departmental, college and university levels.

 Research Interests

Dr. Wang specializes in Silviculture and Forest Ecology. He conducts both applied and foundational ecological research in forest ecosystems, mostly in temperate forests. His current research focuses on how natural and anthropogenic disturbances, including fire, weather extreme, pest/disease, biological invasion, and human activities (including management), affect forest ecosystems and their implication to forest resilience and sustainability.


Refereed Research Articles since 2017 (n = 65; updated July 2, 2021)
(*denote the corresponding author; +denote the leading author completed the whole or a portion of the work in my lab)


Zhang, T., Q.L. Yan*, G.G. Wang, and J.J. Zhu. 2021. The effects of stump size and within-gap position on sprout non-structural carbohydrates concentrations and regeneration in forest gaps vary among species with different shade tolerances. Ecological Processes 10(25), https://doi.org/10.1186/s13717-021-00300-w

Wu+, C., M.D. Ulyshene, C. Shu, Z. Zhang, Y. Zhang, Y. Liu*, and G.G. Wang*. 2021. Stronger effects of termites than microbes on wood decomposition in a subtropical forest. Forest Ecology and Management 493, 119263, https://doi.org/10.1016/j.foreco.2021.119263

Wang+, J., G.G. Wang*, T. Zhang, J. Yuan, L. Yu, J.J. Zhu, and Q.L. Yan*. 2021. Use of direct seeding and seedling planting to restore Korean pine (Pinus koraiensis Sieb. et Zucc.) in secondary forests of Northeast China. Forest Ecology and Management 493, 119243, https://doi.org/10.1016/j.foreco.2021.119243

Lu, D.L., J.J. Zhu, X. Wang, G. Hao, and G.G. Wang. 2021. A systematic evaluation of gap size and within-gap position effects on seedling regeneration in a temperate secondary forest, Northeast China. Forest Ecology and Management 490, 119140, https://doi.org/10.1016/j.foreco.2021.119140.

Xie*+, H.T., Y. Tang, M.K. Yu, and G.G. Wang*. 2021. The effects of afforestation tree species mixing on soil organic carbon stock, nutrients accumulation, and understory vegetation diversity on reclaimed coastal lands in Eastern China. Global Ecology and Conservation 26 e01478. https://doi.org/10.1016/j.gecco.2021.e01478

Zhu, J.J., C.Y. Zhu, D.L. Lu, G.G. Wang, X. Zheng, J.S. Cao, J.X. Zhang. 2021. Regeneration and succession: A 50-year gap dynamic in temperate secondary forests, Northeast China. Forest Ecology and Management 484: 118943 https://doi.org/10.1016/j.foreco.2021.118943

Fang, X.-M. G.G. Wang, Z.-J. Xu, Y.-Y. Zong, X.-L. Zhang, J.-J. Li, H. Wang, F.-S. Chen. 2021. Litter addition and understory removal influenced soil organic carbon quality and mineral nitrogen supply in a subtropical plantation forest. Plant and Soil https://doi.org/10.1007/s11104-020-04787-8


Lu, D.L. J.J. Zhu, D. Wu, Q. Chen, Y. Yu, J. Wang, C. Zhu, H. Liu, T. Gao, G.G. Wang. 2020. Detecting dynamics and variations of crown asymmetry induced by natural gaps in a temperate secondary forest using terrestrial laser scanning. Forest Ecology and Management 473(118289) https://doi.org/10.1016/j.foreco.2020.118289

Lu+ D.L., L.S. Pile, D.P. Yu, J.J. Zhu, D.C. Bragg, G.G. Wang*. 2020. Differential responses of tree species to a severe ice storm and their implications to forest composition in the southeast United States. Forest Ecology and Management 468(118177) https://doi.org/10.1016/j.foreco.2020.118177

Xie+, H.T., J.E. Fawcett, G.G. Wang*. 2020. Fuel dynamics and its implication to fire behavior in loblolly pine-dominated stands after southern pine beetle outbreak. Forest Ecology and Management 466(118130) https://doi.org/10.1016/j.foreco.2020.118130

Wang+, J.P., L. Zhai, J. Ma, J. Zhang*, G.G. Wang*, X. Liu, S. Zhangd, J. Songa, Y. Wu. 2020. Comparative physiological mechanisms of arbuscular mycorrhizal fungi in mitigating salt-induced adverse effects on leaves and roots of Zelkova serrata. Mycorrhiza. 30: 341-355. https://doi.org/10.1007/s00572-020-00954-y

Zhang, P. Y. Wen, L. Wang, H. Zhang, G.G. Wang, T.-G. Wu. 2020. Leaf structural carbohydrate decreased for Pinus thunbergii along a coast-inland gradient. Forests, 11, 449. https://doi.org/10.3390/

Wang+, J., S. Ma, G.G. Wang, L. Xu, Z. Fu, J. Song, J.C. Zhang. 2020. Arbuscular mycorrhizal fungi communities associated with wild plants in a coastal ecosystem. J. For. Res. https://doi.org/10.1007/s11676-020-01127-5

Xiao+, S.S., G.G. Wang, Tang C., Fang, H., Duan J., Yu X.F*. 2020. Effects of One-Year Simulated Nitrogen and Acid Deposition on Soil Respiration in a Subtropical Plantation in China. Forests,11(2): 235. https://doi.org/10.3390/f11020235

Crouch, C.D., B.O. Knapp, S.A. Cohen, M.C. Stambaugh, J.L.Walker, and G.G. Wang. 2020. Site preparation for longleaf pine restoration on hydric sites: Stand development through 15 years after planting. Forest Ecology and Management 461 https://doi.org/10.1016/j.foreco.2020.117928

Wu+, C., Z. Zhang, C. Shua, Q. Mo, H. Wang, F. Kong, Y. Zhang, G.G. Wang*, and Y. Liu*. 2020. The response of coarse woody debris decomposition and microbial community to nutrient additions in a subtropical forest. Forest Ecology and Management 460 https://doi.org/10.1016/j.foreco.2019.117799


Jin+, S.H., B.M. Moule, D. Yu, and G.G. Wang*. 2019. Fire survival of longleaf pine (Pinus palustris) grass stage seedlings: the role of seedling size, root collar position, and resprouting. Forests 10, 1070. https://doi.org/10.3390/f10121070

Wang, J.N., J.Y. Wang, L Wang, H. Zhang, Z. Guo, G.G. Wang, W.K. Smith, T.G. Wu. 2019. Does stoichiometric homeostasis differ among tree organs and with tree age? Forest Ecology and Management 453:117637. https://doi.org/10.1016/j.foreco.2019.117637

Zhang, X., B. Zhang, K. Feeley, G.G. Wang, J. Zhang, and L. Zhai. Ecological contingency in species shifts: Downslope shifts of wood species under warming climate and land-use change. Environmental Research Letters 14: 114033. https://doi.org/10.1088/1748-9326/ab443f

Lu. D., G. Zhang, J.J. Zhu, G.G. Wang, C. Zhu, Q.L. Yan, J. Zhang. 2019. Early natural regeneration patterns of woody species within gaps in a temperate secondary forest. European Journal of Forest Research https://doi.org/10.1007/s10342-019-01219-w

Wang+, F., X.-M. Fang, G.G. Wang*, R. Mao, X.-F. Lin, H. Wang, F.-S. Chen*. 2019. Effects of nutrient addition on foliar phosphorus fractions and their resorption in different-aged leaves of Chinese fir in subtropical China. Plant and Soil https://doi.org/10.1007/s11104-019-04221-8

Wang+, J.P. Z. Zhong, L. Zhu, Y. Yuan, L. Xu, G.G. Wang, L. Zhai, L. Yang, and J. Zhang. 2019. Arbuscular Mycorrhizal fungi effectively enhance the growth of Gleditsia sinensis Lam. seedlings under greenhouse conditions. Forests 10, 567; doi:10.3390/f10070567

Ye, X.M., Y. Zhang, F.S. Chen, G.G. Wang, X.M Fang, X.F. Lin, S.Z. Wan, and P. He. 2019. The effects of simulated deposited nitrogen on nutrient dynamics in decomposing litters across a wide quality spectrum using a 15-N tracing technique. Plant and Soil https://doi.org/10.1007/s11104-019-04158-y

Zhu, C., J.J. Zhu, G.G. Wang, X. Zheng, D. Lu, and T. 2019. Dynamics of gaps and large openings in a secondary forest of Northeast China over 50 years. Annals of Forest Science https://doi.org/10.1007/s13595-019-0844-9

Wang, J.-P., G.G. Wang, B. Zhang, Z. Yuan, Z. Fu, Y. Yuan, L. Zhu, S. Ma, and J. Zhang. 2019. Arbuscular mycorrhizal fungi associated with tree species in a planted forest of Eastern China. Forests 10(5):424 DOI: 10.3390/f10050424

Pile+, L.S., L. Vickers, M. Stambaugh, C. Norman, and G.G. Wang*. 2019. The Tortoise and the Hare: A Race between Native Tree Species and the Invasive Chinese Tallow. Forest Ecology and Management 445: 110-121.

Wu+, C., Z. Zhang, H. Wang, G. Huang, C. Shu, F. Kong, Y. Zhang, G.G. Wang*, and Y. Liu*. 2019. Home-field advantage of CWD decomposition in subtropical forests varied by field sites. Forest Ecology and Management 444: 127-137.

Chen* F.-S., G.G. Wang*, X.-M. Fang, S.-Z. Wan, Y. Zhang, and C. Liang. 2019. Nitrogen deposition effect on forest litter decomposition is interactively regulated by endogenous litter quality and exogenous resource supply. Plant and Soil 437: 413-426.

Wang, F., F.-S. Chen, G.G. Wang, R. Mao, X. Fang, H. Wang, and W. Bu. 2019. Effects of experimental nitrogen addition on nutrients and nonstructural carbohydrates of dominant understory plants in a Chinese fir plantation. Forests 10, 155; doi:10.3390/f10020155

Wu+, C., H. Wang, Q. Mo, Z. Zhang, G. Huang, F. Kong, Y. Liu*, G.G. Wang*. 2019. Effects of elevated UV-B radiation and N deposition on the decomposition of coarse woody debris. The Science of Total Environment 663:170-176


Liu, G., S. Lin, L.S. Pile, F. Zheng, and G.G. Wang. 2018. Effect of potassium permanganate and pyrolysis temperature on the biochar produced from rice straw and suitability of biochars for heavy metal (CD & PB) immobilization in paper sludge. Fresenius Environmental Bulletin 27(12A):9008-9018.

Xie+, H., G.G. Wang*, M. Yu*. 2018. Ecosystem multifunctionality is highly related to the shelterbelt structure and plant species diversity in mixed shelterbelts of eastern China, Global Ecology and Conservation doi:https://doi.org/10.1016/j.gecco.2018.e00470

Liu, G. H. C. Hui, M. Chen, L.S. Pile, G.G. Wang*, F. Wang, P. Shi*. 2018. Variation in individual biomass decreases faster than mean biomass with increasing density of bamboo stands. Journal of Forestry Research 296(11): 245 -7 p. https://doi.org/10.1007/s11676-018-0796-1.

Liu, X., Z. Fu, B. Zhang, L. Zhai, M. Meng, J. Lin, J. Zhuang, G.G. Wang, and J. Zhang. 2018. Effects of sulfuric, nitric, and mixed acid rain on Chinese fir sapling growth in Southern China. Ecotoxicology Environmental Safety 160:154-161. doi: 10.1016/j.ecoenv.2018.04.071. Epub 2018 May 26.

Knapp*, B.O., L.S. Pile, J.L. Walker, and G.G. Wang*. 2018. Fire Effects on a Fire-Adapted Species: Response of Grass-Stage Longleaf Pine Seedlings to Experimental Burning. Fire Ecology 14:2.

Zhang, H., J. Wang, J. Wang, Z. Guo, G.G. Wang, D. Zeng, and T.G. Wu*. 2018. Tree stoichiometry and nutrient resorption along a chronosequence of Metasequoia glyptostroboides forests in coastal China. Forest Ecology and Management 430: 445-450.

Lu+, D., G.G. Wang*, Q.L. Yan, T. Gao, J.J. Zhu*. 2018. Effects of gap size and within-gap position on seedling growth and biomass allocation: Is the gap partitioning hypothesis applicable to the temperate secondary forest ecosystem? Forest Ecology and Management 429: 351-362.

Lu+, D., G.G. Wang*, L.Yu, T. Zhang, J.J. Zhu*. 2018. Seedling survival within forest gaps: the effects of gap size, and within-gap location, and forest type on species of contrasting shade-tolerance. Forestry: An International Journal of Forest Research 91:470-479.

Wu+, C., Q. Mo, H. Wang, Z. Zhang, Q. Ye, G. Huang, Q. Zhou, F. Kong, Y. Liu*, G.G. Wang*. 2018. Moso bamboo (Phyllostachys edulis) expansion affects soil phosphorus cycle in adjacent coniferous forests in subtropical China. Annals of Forest Science 75: 24. https://doi.org/10.1007/s13595-018-0703-0

Wu+, T.G., P. Zhang, L. Zhang, J. Wang, M. Yu, X. Zhou, and G.G. Wang*. 2018. Relationships between shelter effects and optical porosity: a meta-analysis for tree windbreaks. Agricultural and Forest Meteorology 259: 75-81.

Zhang, H., Z.W. Guo, M. Yu, G.G. Wang, and T.G. Wu. 2018. Latitudinal patterns of leaf N, P stoichiometry and nutrient resorption of Metasequoia glyptostroboides along eastern coastline of China. Science of Total Environment 618: 1-6.

Lu+, D., G.G. Wang*, J. Zhang, Y.T. Fang, C. Zhu, J.J. Zhu*. 2018. Converting larch plantations to mixed stands: Effects of canopy treatment on the survival and growth of planted seedlings with contrasting shade tolerance. Forest Ecology and Management 409:19-28.

Shearman+, T.M., G.G. Wang*, R.K. Peet, T.R. Wentworth, M.P. Schafale, and A.S. Weakley. 2018. A community analysis for forest ecosystems associated with Persea borbonia and P. palustris, two species at risk from Laurel Wilt Disease in the southeastern United States. Castanea 83:3-27.


Yang, X., P. Zhang, Q. Wu, H. Yan, M. Yu, G.G. Wang, and T.G. Wu. 2017 Effects of Simulated Wind Load on Leaf Photosynthesis and Carbohydrate Allocation in Eight Quercus Species. Journal of Biobased Materials and Bioenergy 11: 652-657.

Pile+, L.S., G.G. Wang*, B.O. Knapp, G. Liu, and D. Yu. 2017. Morphological and physiological adaptations to frequent surface fire regimes: A comparative study on seedlings of three Pinus species native to southeast United States. Annals of Forest Science 74(4): 68-80.

Coates, T.A., Alex T. Chow, D.L. Hagan, T.A. Waldrop, G.G. Wang, W.C. Bridges, Jr., M.-F. Rogers, and J.H. Dozier. 2017. Thermocouple probe orientation affects prescribed fire behavior estimation. Journal of Environmental 47(1):170-176 DOI10.2134/jeq2017.02.0055

Coates, T.A., A.T. Chow, D.L. Hagan, G.G. Wang, W.C. Bridges, and J.H. Dozier. 2017. Frequent Prescribed Burning as a Long-term Practice in Longleaf Pine Forests Does Not Affect Detrital Chemical Composition. Journal of Environmental Quality 46:1020-1027.

Pile+, L.S., G.G. Wang*, J.P. Stovall, G.S. Wheeler, and E. Siemann. 2017. Mechanisms of Chinese tallow (Triadica sebifera) invasion and their management implications – A review. Forest Ecology and Management 404: 1-13.

Pile+, L.S., G.G. Wang*, T.A. Waldrop, J.L. Walker, W.C. Bridges, and P.A. Layton. 2017. Developing an ecologically-based integrated management approach to control Chinese tallow (Triadica sebifera) invasion in maritime forests. Journal of Forestry 115(6):522-529.

Cheng+, X., M. Yu, and G.G. Wang+. 2017. Effects of Thinning on Soil Organic Carbon Fractions and Soil Properties in Cunninghamia lanceolata Stands in Eastern China. Forests 8(6), 198; doi:10.3390/f8060198

Pile+, L.S., G.G. Wang*, B.O. Knapp, J.L. Walker, M.C. Stambaugh. 2017. Chinese tallow (Triadica sebifera) invasion in maritime forests: the role of anthropogenic disturbances and its management implication. Forest Ecology and Management 398: 10-24.

Hu+, H., G.G. Wang*, B.L. Bauerle, and R.J. Klos. 2017. Drought impact on forest regeneration in the Southeast USA. Ecosphere 8(4): e01772.

Zhang, H., X. Yang, J. Wang, G.G. Wang, M. Yu, and T.G. Wu*. 2017. Leaf N and P stoichiometry in relation to leaf shape and plant size for Quercus acutissima provenances across China. Scientific Reports 7: 46133.


Google Scholar
Research Gate