G. Geoff Wang

Forestry and Environmental Conservation Department

Office: 226 Lehotsky Hall
Phone: 864-656-4864
Fax: 864-656-3304

Email: gwang@clemson.edu

 Educational Background

Ph.D. Forest Ecology
Univ. of British Columbia

 Courses Taught

Silviculture Laboratory
Fire Ecology and Management
Fire Ecology and Management Laboratory
Creative Inquiry (Undergraduate Research)
Selected Topics in Forest Resources
Master Thesis Research
Doctoral Dissertation Research


Dr. Wang directs the Silviculture and Ecology Lab (SEL) at Clemson University. He has been either PI/ Project Leader or Co-PI for ~50 externally funded projects. Currently, Dr. Wang has eight ongoing research projects, including an active program of international research collaboration. He has published ~130 peer-reviewed journal articles and over ~70 non-refereed publications. He has also delivered ~250 conference presentations, including ~70 invited seminars. His research has made a great impact in the field as indicated by ~2700 citations, an h-index of 28, and an RG score of 38.

Dr. Wang has taught multiple undergraduate graduate courses and mentored 21 post-doctoral scientists, 14 doctoral students, 22 master students, and seven honors students. Students under his mentorship have won many prestigious scholarships and awards. Most Ph.D. students and post-docs trained in his lab currently hold tenure-track faculty positions or equivalent.

Dr. Wang currently serves as an editor for Forestry: An International Journal of Forestry Research; an Associates Editor for New Forests, Canadian Journal of Forest Research, and Journal of Forestry Research; as a member in the editorial board for Journal of Forest and Environment. Dr. Wang has been frequently consulted nationally and internationally by scientific journals, grant agencies, government agencies, professional forestry associations, and industries. Internally, he has also served and chaired committees at departmental, college and university levels.

 Research Interests

Dr. Wang specializes in Silviculture and Forest Ecology. He conducts both applied and foundational ecological research in forest ecosystems, mostly in temperate forests. His current research focuses on how natural and anthropogenic disturbances, including fire, weather extreme, pest/disease, biological invasion, and human activities (including management), affect forest ecosystems and their implication to forest resilience and sustainability.


Refereed Research Articles since 2015 (n = 55; updated Sept 2019)
(*denote the corresponding author; +denote the leading author completed the work in the lab)

Lu. D., G. Zhang, J.J. Zhu, G.G. Wang, C. Zhu, Q.L. Yan, J. Zhang. 2019. Early natural regeneration patterns of woody species within gaps in a temperate secondary forest. European Journal of Forest Research https://doi.org/10.1007/s10342-019-01219-w

Wang, F., X.-M. Fang, G.G. Wang*, R. Mao, X.-F. Lin, H. Wang, F.-S. Chen*. 2019. Effects of nutrient addition on foliar phosphorus fractions and their resorption in different-aged leaves of Chinese fir in subtropical China. Plant and Soil https://doi.org/10.1007/s11104-019-04221-8

Wang J.P. Z. Zhong, L. Zhu, Y. Yuan, L. Xu, G.G. Wang, L. Zhai, L. Yang, and J. Zhang. 2019. Arbuscular Mycorrhizal fungi effectively enhance the growth of Gleditsia sinensis Lam. seedlings under greenhouse conditions. Forests 10, 567; doi:10.3390/f10070567

Ye, X.M., Y. Zhang, F.S. Chen, G.G. Wang, X.M Fang, X.F. Lin, S.Z. Wan, and P. He. 2019. The effects of simulated deposited nitrogen on nutrient dynamics in decomposing litters across a wide quality spectrum using a 15-N tracing technique. Plant and Soil https://doi.org/10.1007/s11104-019-04158-y

Zhu, C., J.J. Zhu, G.G. Wang, X. Zheng, D. Lu, and T. 2019. Dynamics of gaps and large openings in a secondary forest of Northeast China over 50 years. Annals of Forest Science https://doi.org/10.1007/s13595-019-0844-9

Wang, J.-P., G.G. Wang, B. Zhang, Z. Yuan, Z. Fu, Y. Yuan, L. Zhu, S. Ma, and J. Zhang. 2019. Arbuscular mycorrhizal fungi associated with tree species in a planted forest of Eastern China. Forests 10(5):424 DOI: 10.3390/f10050424

Pile, L.S., L. Vickers, M. Stambaugh, C. Norman, and G.G. Wang*. 2019. The Tortoise and the Hare: A Race between Native Tree Species and the Invasive Chinese Tallow. Forest Ecology and Management 445: 110-121.

Wu, C., Z. Zhang, H. Wang, G. Huang, C. Shu, F. Kong, Y. Zhang, G.G. Wang*, and Y. Liu*. 2019. Home-field advantage of CWD decomposition in subtropical forests varied by field sites. Forest Ecology and Management 444: 127-137.

Chen* F.-S., G.G. Wang*, X.-M. Fang, S.-Z. Wan, Y. Zhang, and C. Liang. 2019. Nitrogen deposition effect on forest litter decomposition is interactively regulated by endogenous litter quality and exogenous resource supply. Plant and Soil 437: 413-426.

Wang, F., F.-S. Chen, G.G. Wang, R. Mao, X. Fang, H. Wang, and W. Bu. 2019. Effects of experimental nitrogen addition on nutrients and nonstructural carbohydrates of dominant understory plants in a Chinese fir plantation. Forests 10, 155; doi:10.3390/f10020155

Wu, C., H. Wang, Q. Mo, Z. Zhang, G. Huang, F. Kong, Y. Liu*, G.G. Wang*. 2019. Effects of elevated UV-B radiation and N deposition on the decomposition of coarse woody debris. The Science of Total Environment 663:170-176


Liu, G., S. Lin, L.S. Pile, F. Zheng, and G.G. Wang. 2018. Effect of potassium permanganate and pyrolysis temperature on the biochar produced from rice straw and suitability of biochars for heavy metal (CD & PB) immobilization in paper sludge. Fresenius Environmental Bulletin 27(12A):9008-9018.

Xie, H., G.G. Wang*, M. Yu*. 2018. Ecosystem multifunctionality is highly related to the shelterbelt structure and plant species diversity in mixed shelterbelts of eastern China, Global Ecology and Conservation doi:https://doi.org/10.1016/j.gecco.2018.e00470

Liu, G. H. C. Hui, M. Chen, L.S. Pile, G.G. Wang*, F. Wang, P. Shi*. 2018. Variation in individual biomass decreases faster than mean biomass with increasing density of bamboo stands. Journal of Forestry Research 296(11): 245 -7 p. https://doi.org/10.1007/s11676-018-0796-1.

Liu, X., Z. Fu, B. Zhang, L. Zhai, M. Meng, J. Lin, J. Zhuang, G.G. Wang, and J. Zhang. 2018. Effects of sulfuric, nitric, and mixed acid rain on Chinese fir sapling growth in Southern China. Ecotoxicology Environmental Safety 160:154-161. doi: 10.1016/j.ecoenv.2018.04.071. Epub 2018 May 26.

Knapp*, B.O., L.S. Pile, J.L. Walker, and G.G. Wang*. 2018. Fire Effects on a Fire-Adapted Species: Response of Grass-Stage Longleaf Pine Seedlings to Experimental Burning. Fire Ecology 14:2.

Zhang, H., J. Wang, J. Wang, Z. Guo, G.G. Wang, D. Zeng, and T.G. Wu*. 2018. Tree stoichiometry and nutrient resorption along a chronosequence of Metasequoia glyptostroboides forests in coastal China. Forest Ecology and Management 430: 445-450.

Lu, D., G.G. Wang*, Q.L. Yan, T. Gao, J.J. Zhu*. 2018. Effects of gap size and within-gap position on seedling growth and biomass allocation: Is the gap partitioning hypothesis applicable to the temperate secondary forest ecosystem? Forest Ecology and Management 429: 351-362

Lu, D., G.G. Wang*, L.Yu, T. Zhang, J.J. Zhu*. 2018. Seedling survival within forest gaps: the effects of gap size, and within-gap location, and forest type on species of contrasting shade-tolerance. Forestry: An International Journal of Forest Research 91:470-479.

Wu, C., Q. Mo, H. Wang, Z. Zhang, Q. Ye, G. Huang, Q. Zhou, F. Kong, Y. Liu*, G.G. Wang*. 2018. Moso bamboo (Phyllostachys edulis) expansion affects soil phosphorus cycle in adjacent coniferous forests in subtropical China. Annals of Forest Science 75: 24. https://doi.org/10.1007/s13595-018-0703-0

Wu, T.G., P. Zhang, L. Zhang, J. Wang, M. Yu, X. Zhou, and G.G. Wang*. 2018. Relationships between shelter effects and optical porosity: a meta-analysis for tree windbreaks. Agricultural and Forest Meteorology 259: 75-81.

Zhang, H., Z.W. Guo, M. Yu, G.G. Wang, and T.G. Wu. 2018. Latitudinal patterns of leaf N, P stoichiometry and nutrient resorption of Metasequoia glyptostroboides along eastern coastline of China. Science of Total Environment 618: 1-6.

Lu, D., G.G. Wang*, J. Zhang, Y.T. Fang, C. Zhu, J.J. Zhu*. 2018. Converting larch plantations to mixed stands: Effects of canopy treatment on the survival and growth of planted seedlings with contrasting shade tolerance. Forest Ecology and Management 409:19-28.

Shearman, T.M., G.G. Wang*, R.K. Peet, T.R. Wentworth, M.P. Schafale, and A.S. Weakley. 2018. A community analysis for forest ecosystems associated with Persea borbonia and P. palustris, two species at risk from Laurel Wilt Disease in the southeastern United States. Castanea 83:3-27.


Yang, X., P. Zhang, Q. Wu, H. Yan, M. Yu, G.G. Wang, and T.G. Wu. 2017 Effects of Simulated Wind Load on Leaf Photosynthesis and Carbohydrate Allocation in Eight Quercus Species. Journal of Biobased Materials and Bioenergy 11: 652–657.

Pile, L.S., G.G. Wang*, B.O. Knapp, G. Liu, and D. Yu. 2017. Morphological and physiological adaptations to frequent surface fire regimes: A comparative study on seedlings of three Pinus species native to southeast United States. Annals of Forest Science 74(4): 68-80.

Coates, T.A., Alex T. Chow, D.L. Hagan, T.A. Waldrop, G.G. Wang, W.C. Bridges, Jr., M.-F. Rogers, and J.H. Dozier. 2017. Thermocouple probe orientation affects prescribed fire behavior estimation. Journal of Environmental 47(1):170-176 DOI10.2134/jeq2017.02.0055

Coates, T.A., A.T. Chow, D.L. Hagan, G.G. Wang, W.C. Bridges, and J.H. Dozier. 2017. Frequent Prescribed Burning as a Long-term Practice in Longleaf Pine Forests Does Not Affect Detrital Chemical Composition. Journal of Environmental Quality 46:1020-1027.

Pile, L.S., G.G. Wang*, J.P. Stovall, G.S. Wheeler, and E. Siemann. 2017. Mechanisms of Chinese tallow (Triadica sebifera) invasion and their management implications – A review. Forest Ecology and Management 404: 1-13.

Pile, L.S., G.G. Wang*, T.A. Waldrop, J.L. Walker, W.C. Bridges, and P.A. Layton. 2017. Developing an ecologically-based integrated management approach to control Chinese tallow (Triadica sebifera) invasion in maritime forests. Journal of Forestry 115(6):522-529.

Cheng, X., M. Yu, and G.G. Wang+. 2017. Effects of Thinning on Soil Organic Carbon Fractions and Soil Properties in Cunninghamia lanceolata Stands in Eastern China. Forests
8(6), 198; doi:10.3390/f8060198

Pile, L.S., G.G. Wang*, B.O. Knapp, J.L. Walker, M.C. Stambaugh. 2017. Chinese tallow (Triadica sebifera) invasion in maritime forests: the role of anthropogenic disturbances and its management implication. Forest Ecology and Management 398: 10-24.

Hu, H., G.G. Wang*, B.L. Bauerle, and R.J. Klos. 2017. Drought impact on forest regeneration in the Southeast USA. Ecosphere 8(4): e01772.

Zhang, H., X. Yang, J. Wang, G.G. Wang, M. Yu, and T.G. Wu*. 2017. Leaf N and P stoichiometry in relation to leaf shape and plant size for Quercus acutissima provenances across China. Scientific Reports 7: 46133.


Yan, Q.L., J.J. Zhu; Q. Gang, L. Huang, and G.G. Wang. 2016. Comparison of Spatial Distribution Patterns of Seed Rain between Larch Plantations and Adjacent Secondary Forests in Northeast China. Forest Science 62(6): 652-662; DOI: http://dx.doi.org/10.5849/forsci.15-048

Chhin, S. and G.G. Wang. 2016. Climatic Sensitivity of a Mixed Forest Association of White Spruce and Trembling Aspen at Their Southern Range Limit. Forests 7(10): 235; doi:10.3390/f7100235. http://www.mdpi.com/1999-4907/7/10/235

Wu, T.G., P. Zhang, L. Zhang, G.G. Wang, and M.K. Yu. 2016. Morphological response of eight Quercus species to simulated wind load. PLOS ONE 11(9): e0163613. doi: 10.1371/journal.pone.0163613.

Yueqin Cheng, Jingmian Duan, Zhenbin Jiao, G. Geoff Wang, Fengming Yan, and Hongwei Wang*. 2016. Cytoplasmic DNA disclose high nucleotide diversity and different phylogenetic pattern in Taihangia rupestris. Biochemical Systematics and Ecology 66: 201-208.

Ruan, Xiao, Cunde Pan, Run Liu, Zhaohui, Li, Shuling Li; De-An Jiang, Jing-Chi Zhang, G. Geoff Wang, Yin-Xian Zhao, and Qiang Wang. 2016. Effects of climate warming on plant autotoxicity in forest evolution -a case simulation analysis for Picea schrenkiana regeneration. Ecology and Evolution doi:10.1002/ece3.2315

Wu, T.G., H. Zhang, G.G. Wang, and M.K. Yu. 2016. Effects of environment and genetics on leaf N and P stoichiometry for 29 Quercus acutissima provenances in China. European Journal of Forest Research 135(4): 795-802.

Yan, Y., Q. Zhang, G.G. Wang*, and Y.-M. Fang*. 2016. Atmospheric Deposition of Heavy Metals in Wuxi, China: Estimation Based on Native Moss Analysis. Environmental Monitoring and Assessment. 188(6):360. doi: 10.1007/s10661-016-5315-2

Knapp*, B.O., G.G. Wang*, J.L. Walker, and H. Hu. 2016. Using silvicultural practices to regulate competition, resource availability, and growing conditions for Pinus palustris seedlings underplanted in Pinus taeda forests. Canadian Journal of Forest Research 46: 902-913

Liu, G., Q. Xu, X. Dong, J. Yang, L.S. Pile, G.G. Wang, and F. Wang. 2016. Effect of protective gas and pyrolysis temperature on the biochar produced from three plants of Gramineae: physical and chemical characterization. Waste Biomass Valorization. DOI: 10.1007/s12649-016-9534-0

Hu, H., B.O. Knapp, G.G. Wang*, and J.L. Walker. 2016. Silvicultural treatments for converting loblolly pine to longleaf pine dominance: Effects on ground layer and midstory vegetation. Journal of Applied Vegetation Science 19: 280-290

Liu, G., P. Shi, Q. Xu, X. Dong, F. Wang, G.G. Wang*, and C. Hui. 2016. Does the size-density relationship developed for bamboo species conform to the self-thinning rule? For. Ecol. Manage. 361: 339-345.

Pile, L.S., C.A. Maier, G.G. Wang*, D. Yu, T.M. Shearman. 2016. Responses of two genetically superior loblolly pine clonal ideotypes to a severe ice storm. Forest Ecology and Management 360: 213-220.


Jiao, Z., Y. Cheng, H. Wang, C. Lei, G.G. Wang, L. Han. 2015. Isolation and Characterization of Microsatellite Loci in Rehmannia glutinosa (Scrophulariaceae), a Medicinal Herb. Applications in Plant Sciences 3(10): apps1500054

Zhang, J.Y., L.J. Lin, L. Yang, X. Ruan, X.J. Li, C.D. Pan, H. Zhang, G.G. Wang, and Q. Wang. 2015. Comparative allelopathic potentials of American chestnut (Castanea dentata) and Chinese chestnut (C. mollissima). Allelopathy Journal 36: 303-314.

Addington, R.N., B.O. Knapp, G.G. Sorrell, M.L. Elmore, G.G. Wang, and J.L. Walker. 2015. Factors affecting broadleaf woody vegetation in upland pine forests managed for longleaf pine restoration. Forest Ecology and Management 354:130-138

Zhu, J., G. Zhang, G.G. Wang, Q. Yan, D. Lu, X. Li and X. Zheng. 2015. On the size of forest gaps: can its lower and upper limits be objectively defined? Agricultural and Forestry Meteorology 213: 64-76.

Zhang, B., X. Liu, D.L. DeAngelis, W.-M. Ni, and G.G. Wang. 2015. Effects of dispersal on total biomass in a patchy, heterogeneous system: Analysis and experiment. Mathematical Biosciences 264: 54-62.

Stottlemyer, Aaron D., T.A. Waldrop, and G.G. Wang+. 2015. Surface Fuel Characteristics in Beetle-killed Southern Pine Stands and Impacts of Prescribed Fire and Mastication as Fuel Reduction Treatments. Ecological Engineering 81: 514-524.

Pile. L.S., G.G. Wang*, R. Polomski, G. Yarrow and C.M. Stuyck. 2015. Potential for non-native endozoochorous seed dispersal by white-tailed deer in a southeastern maritime forest. Invasive Plant Management and Science 8:32-43.

Jin, S.H., X.Q. Li, G.G. Wang+, and X.T. Zhu. 2015. Brassinosteroids alleviate high temperature injury in Ficus concinna seedlings via maintaining higher antioxidant defense and glyoxalase systems. AoB Plants AoB PLANTS7: plv009; doi:10.1093/aobpla/plv009

Shearman, TM, G.G. Wang*, and WC Bridges. 2015. Impacts of laurel wilt disease on redbay (Persea borbonia): An assessment based on forest inventory and analysis data. Biological Invasions 17: 1371-1382