Patrick Jodice

Professor, Leader SC Cooperative Research Unit
Leader, SC Cooperative Research Unit
Forestry and Environmental Conservation Department

Office: 238 Lehotsky
Phone: 864-656-6190
Vita: Download CV


 Educational Background

Ph.D. Wildlife Science
Oregon State University 1999

M.S. Wildlife Ecology & Conservation
University of Florida 1990

B.S. Wildlife Management
University of Maine 1983

 Courses Taught

Conservation Physiology
Foundations of Ecology
Wildlife Energetics
Seabird Ecology


I am a Wildlife Research Biologist with the USGS South Carolina Cooperative Fish and Wildlife Research Unit. As such I focus primarily on graduate education and research through graduate students. My current research is focused primarily on the ecology and conservation of seabirds and shorebirds. I currently serve as the Chair of the World Seabird Union.

 Research Interests

My research is focused on conservation and ecology of seabirds and shorebirds, and wildlife conservation physiology. I have conducted research and monitoring programs in a variety of marine and terrestrial ecosystems and have studied both avian and mammalian populations. All of my research has been driven by management and conservation issues, although the results are often of both an applied and basic nature.
My current and prior work has emphasized wildlife conservation with emphases in marine ornithology, avian ecology, reproductive energetics, foraging ecology, survey design and methodology, and urban ecology.


* = graduate student advisee

Jodice, P.G.R., P.E. Michael, J.S. Gleason, J.C. Haney, Y.G. Satgé. 2021. Revising the marine range of the endangered black-capped petrel Pterodroma hasitata: Occurrence in the northern Gulf of Mexico and exposure to conservation threats. Endangered Species Research 46: 49-65.

Streker, R.A.*, J.S. Lamb*, J. Dindo, P.G.R. Jodice. 2021. Fine-scale weather patterns drive reproductive success in the Brown Pelican. Waterbirds 44:153-166.

Wilkinson, B.P.*, A.R. Robuck, R. Lohmann, H. Pickard, P.G.R. Jodice. 2021. Urban proximity while breeding is not a predictor of perfluoroalkyl substance contamination in the eggs of brown pelicans. Science of the Total Environment.

Takahashi, F., F. Sanders, P.G.R. Jodice. 2021. Spatial and temporal overlap between foraging shorebirds and spawning Horseshoe Crabs (Limulus polyphemus) in the Cape Romain - Santee Delta Region of the U.S. Atlantic Coast. Wilson Journal of Ornithology 133:58-72.

Fiorello, C., P.G.R. Jodice, J.S. Lamb, Y.G. Satge, K. Mills, M. Zicardi. 2021. Post-release survival of California Brown Pelicans following oiling and rehabilitation after the Refugio oil spill. Journal of Wildlife Diseases 57(3). DOI: 10.7589/JWD-D-20-00171

Satgé, Y.G.^, E. Rupp, A. Brown, and P.G.R Jodice. 2020. Habitat modelling locates nesting areas of the Endangered Black-capped Petrel Pterodroma hasitata on Hispaniola and identifies habitat loss. Bird Conservation International 1-18. doi:10.1017/S0959270920000490.

Fallon, J.A., E.P. Smith. N. Shoch, J.D. Paruk, E.A. Adams, D.C. Evers, P.G.R. Jodice, W.A. Hopkins. 2020. Ultraviolet-assisted oiling assessment improves detection of oiled birds experiencing clinical signs of hemolytic anemia exposed to the Deepwater Horizon oil spill. Ecotoxicology.

Ramos, R., V.H. Paiva, Z. Zajkova, C. Precheur, A.I. Fagundes, P.G.R. Jodice, W. Mackin, V. Neves, F. Zino, V. Bretagnolle, J. Gonzalez-Solis. 2020. Spatial ecology of closely related taxa: the case of the little shearwater complex in the North Atlantic Ocean. Zoological Journal of the Linnean Society XX 1-21.

Wilkinson, B.P.*, A.M. Haynes-Sutton, L. Meggs, and P.G.R Jodice. Accepted. High spatial fidelity among foraging trips of Masked Boobies from Pedro Bank, Jamaica. PLoOS ONE 15(4): e0231654.

Lamb, J.S.*, Y.G. Satgé^, R.A. Streker*, P.G.R Jodice. Ecological drivers of Brown Pelican movement patterns, health, and reproductive success in the Gulf of Mexico. U.S. Dept. of the Interior, Bureau of Ocean Energy Management, Gulf of Mexico OCS Region, New Orleans, LA. OCS Study BOEM-2020-036. 232 pp.

Ndu, U., J.S. Lamb, S. Janssen, R. Rossi, Y. Satgé, and P.G.R Jodice. 2020. Mercury, cadmium, copper, arsenic, and selenium measurements in the feathers of adult Eastern Brown Pelicans and chicks in multiple breeding grounds in the northern Gulf of Mexico. Environmental Research Letters 192, 286.

Zaluski, S., L.M. Soanes, J.A. Bright, A. George, P.G.R. Jodice, K. Meyer, N. Woodfield-Pascoe, J.A. Green. 2020. Potential threats facing a globally important population of the Magnificent Frigatebird. Tropical Zoology 32: 188-201. /doi/full/10.1080/03946975.2019.1682352.

Lamb, J.S.*, Y.G. Satgé^, P.G.R. Jodice. 2020. Seasonal variation in environmental and behavioral drivers of annual-cycle habitat selection in a nearshore seabird. Diversity and Distributions.

Wilkinson, B.P.*, Y.G. Satgé^, J.S. Lamb, P.G.R. Jodice. 2019. Tropical cyclones alter short-term activity patterns of a coastal seabird. Movement Ecology 7:30.

Lawson, A.J.*, C.T. Moore, T.R. Rainwater, F.M. Nilsen, P.M. Wilkinson, R.H. Lowers, L. J. Guillette Jr., K.W. McFadden, and P.G.R. Jodice. 2019. Non-linear patterns in mercury bioaccumulation patterns in American alligators are a function of predicted age. Science of the Total Environment. https://doi.oprg/10.1016/j.scitotenv.2019.135103

Jodice, P.G.R., E.M. Adams, J.S. Lamb, Y. Satgé^, J.S. Gleason. 2019. Strategic Bird Monitoring Guidelines for the Northern Gulf of Mexico: Seabirds. Pages 133 – 172. In Wilson et al. (eds.). Strategic Bird Monitoring Guidelines for the Northern Gulf of Mexico, Mississippi Agricultural and Forestry Extension Research Bulletin 1228, Mississippi State University.

Ottinger, M.A., T. Maness, J.K. Grace, R. Wilson, P.G.R. Jodice. 2019. Strategic Avian Monitoring Plan for the Gulf of Mexico: Avian Health Assessments. In Wilson et al. (eds.). Strategic Bird Monitoring Guidelines for the Northern Gulf of Mexico, Mississippi Agricultural and Forestry Extension Research Bulletin 1228, Mississippi State University.

Ferguson, L.M., Satgé^, J. Tavano^, and P.G.R. Jodice. 2018. Seabird colony registry and atlas for the Southeastern U.S. Final Report for U.S. Fish and Wildlife Service. South Carolina Cooperative Fish and Wildlife Research Unit, Clemson, SC.

Lamb, J.S.*, C.V. Fiorello, Y.G. Satgé^, K. Mills-Parker, M. Ziccardi, P.G.R. Jodice. 2018. Movement patterns of California brown pelicans following oiling and rehabilitation. Marine Pollution Bulletin 131:22-31.

Loeb, S.C., P.G.R. Jodice. 2018. Activity of southeastern bats along sandstone cliffs used for rock climbing. Journal of Fish and Wildlife Management 9:255-265; e1944-687x.

Lamb, J.S.*, D.J. Newstead, L.M. Koczur, B.M. Ballard, C.M. Green, P.G.R. Jodice. 2018. A bridge between oceans: Overland migration of marine birds in a wind energy corridor. Journal of Avian Biology e01474 doi: 10.1111/jav.01474

Fallon, J.A., E.P. Smith. N. Shoch, J.D. Paruk, E.A. Adams, D.C. Evers, P.G.R. Jodice, C. Perkins, S. Shulte, W.A. Hopkins. Accepted. Hematological indices of physiological injury to lightly oiled birds from the Deepwater Horizon oil spill. Environmental Toxicology and Chemistry.

Lamb, J.S.*, Y.G. Satge^, P.G.R. Jodice. 2017. Diet composition and provisioning rates of nestlings determine reproductive success in a subtropical seabird. Marine Ecology Progress Series 581:149-164.

Lamb, J.S.*, Y.G. Satge^, P.G.R. Jodice. 2017. Influence of density-dependent competition on foraging and migratory behavior of a subtropical colonial seabird. Ecology and Evolution 2017;00:1–13.

Poli, C.L.*, A-L. Harrison, A. Vallarino, P.D. Gerard, Jodice, P.G.R. 2017. Dynamic oceanography predicts fine scale foraging behavior of Masked Boobies in the Gulf of Mexico. PLoS ONE 12(6): e0178318.

Zinsser, E.*, F.J. Sanders, P. Gerard, P.G.R. Jodice. 2017. Daily survival rate and habitat characteristics of nests of Wilson’s Plover. Southeastern Naturalist 16:149-156.

Ramos, R., N. Carlile, J. Madeiros, I. Ramirez, V. Paiva, H. Dinis, F. Zino, M. Biscoito, G. Leal, L. Bugoni, P.G.R. Jodice, P.G. Ryan, J. Gonzalez-Solis. 2017. It is the time for oceanic seabirds: tracking year-round distribution of gadfly petrels across the Atlantic Ocean. Diversity and Distributions.

Haney, J.C., P.G.R. Jodice, W.A. Montevecchi, D.C. Evers. 2017. Challenges to oil spill assessment for seabirds in the deep ocean. Archives of Environmental Contamination and Toxicology. doi:10.1007/s00244-016-0355-8

Lamb, J.S.*, K.M. O’Reilly, P.G.R. Jodice. 2016. Physical condition and stress levels during early development reflect feeding rates and predict pre- and post-fledging survival in a nearshore seabird. Conservation Physiology 4 (1): cow060. doi: 10.1093/conphys/cow060

Lamb, J.S.*, Y.G. Satge^, C.V. Fiorello, P.G.R. Jodice. 2016. Behavioral and reproductive effects of bird-borne data logger attachment on Brown Pelicans on three temporal scales. Journal of Ornithology. doi:10.1007/s10336-016-1418-3

Collins, S.A.*, F.J. Sanders, P.G.R. Jodice. 2016. Assessing conservation tools for an at-risk shorebird: Feasibility of headstarting for American Oystercatchers. Bird Conservation International 26: 451-465.

Jodice, P.G.R., R.A. Ronconi, E. Rupp, G.E. Wallace, Y. Satge. 2015. First satellite tracks of the endangered Black-capped Petrel. Endangered Species Research 29:23-33. doi: 10.3354/esr00697

Ferguson, L.M.*, T.M. Norton, C. Cray, M. Olivia, P.G.R. Jodice. 2014. Health assessments of brown pelican nestlings from colonies in South Carolina and Georgia. Journal of Zoo and Wildlife Medicine 45:835-845.

Jodice, P.G.R., J.M. Thibault*, S.A. Collins*, M. Spinks, F.J. Sanders. 2014. Reproductive ecology of American Oystercatchers nesting on shell rakes. Condor 116:588-598.

Lucas, J.S.*, S.C. Loeb, P.G.R. Jodice. 2015. Roost selection by Rafinesque’s big-eared bats (Corynorhinus rafinesquii) in optimal habitat. Acta Chiropterologica 17:131-141.

Brooks, G.L.*, F.J. Sanders, P.D. Gerard, and P.G.R. Jodice. 2014. Daily survival rates for nests of Black Skimmers from a core breeding area of the Southeastern USA. Wilson Journal of Ornithology 126:443-450.

Carlson-Bremer, D., T.M. Norton, F.J. Sanders, B. Winn, M. Spinks, B.A. Glatt, L. Mazzaro, P.G.R. Jodice, T. Chen, E.S. Dierenfeld. 2014. Circulating fat-soluble vitamin concentrations and nutrient composition of aquatic prey eaten by American Oystercatchers in the southeastern United States. Journal of Avian Medicine and Surgery 28:216-224.

Hackney, A., R. Baldwin, P.G.R. Jodice. 2013. Mapping risk for nest predation on a barrier island. Journal of Coastal Conservation 17:615-621.

Giles, M.M.*, R.F. Baldwin, J. Stanton, M. Epstein, P.G.R. Jodice. 2013. Spring migratory pathways and migration chronology of Canada Geese wintering at the Santee NWR. Canadian Field Naturalist 127: 17-25.

Brooks, G.L.*, F.J. Sanders, P.D. Gerard, P.G.R. Jodice. 2013. Daily survival rate for nests and chicks of Least Terns (Sternula antillarum). Waterbirds 36:1-10.

Jodice, P.G.R., J. Tavano, W. Mackin. 2013. Chapter 8: Marine and coastal birds and bats. Pages 475-587 In: Michel, J. (ed.). South Atlantic information resources: data search and literature synthesis. U.S. Department of the Interior, Bureau of Ocean Energy Management, Regulation, and Enforcement, Gulf of Mexico OCS Region, New Orleans, LA. OCS Study BOEM 2013-01157.

Vander Pol, S.S., D.W. Anderson, P.G.R. Jodice, J.E. Stuckey. 2012. East versus west: organic contaminant differences in Brown Pelican eggs from South Carolina, USA and the Gulf of California, Mexico. Science and the Total Environment 438: 527-532.

Jodice, P.G.R., L.C. Wickliffe*, E.B. Sachs*. 2011. Seabird use of discards from a nearshore shrimp fishery in the South Atlantic Bight, USA. Marine Biology 158:2289-2298.

Jodice, P.G.R., R.M. Suryan. 2010. The transboundary nature of seabird ecology. Pages 139-165 In Landscape Scale Conservation Planning (S. Trombulak and R. Baldwin, eds.). Springer.

Hand, C.*, F. Sanders, P.G.R. Jodice. 2010. Foraging proficiency during the nonbreeding season in a specialized forager: Are juvenile American Oystercatchers (Haematopus palliatus) 'bumble-beaks'* compared to adults? Condor 112:670-675.

Thibault, J.*, F. Sanders, P.G.R. Jodice. 2010. Parental attendance and brood success in American Oystercatchers. Waterbirds 33:511-517.

Wickliffe, L.C.*, P.G.R. Jodice. 2010. Abundance of nearshore seabirds at shrimp trawlers in South Carolina. Marine Ornithology 38:31-39.

Meehan, K.*, P.G.R. Jodice. 2010. Landscape scale correlates of fox squirrel (Sciurus niger) presence on golf courses in coastal South Carolina. Southeastern Naturalist 9:573-586.

Eggert, L.M.F. *, P.G.R. Jodice, K.M. O’Reilly. 2010. Stress response of Brown Pelican nestlings to ectoparasite infestation. General and Comparative Endocrinology. 166: 33-38.

Sachs, E.B.*, P.G.R. Jodice. 2009. Behavior of parent and nestling Brown Pelicans during early brood-rearing. Waterbirds 32:276-281.

Peden-Adams, M.M., J.E. Stuckey, K. Gaworecki, J. Berger-Ritchie, K. Bryant, P.G.R. Jodice, T.R. Scott, S. Boone, W.D. McGuinn, J.C. DeWitt, D.E. Keil. 2009. Developmental toxicity in white leghorn chickens following in ovo exposure of perfluorooctane sulfonate (PFOS). Reproductive Toxicology 27:307-318.

Friebel, B.A.*, P.G.R. Jodice. 2009. Home range and habitat use of wild hogs in the Congaree National Park, South Carolina. Human-Wildlife Conflicts 3:49-63.
Jodice, P.G.R., D.D. Roby, K.R. Turco, R.M. Suryan, D.B. Irons, J.F. Piatt, M.T. Shultz, D.G. Roseneau, A.B. Kettle. 2008. Growth rates of Black- legged Kittiwake Rissa tridactyla chicks in relation to delivery rate, size and energy density of meals. Marine Ornithology 36:107-114.

Eggert, L.M.F.*, P.G.R. Jodice. 2008. Growth of Brown Pelican nestlings exposed to sublethal levels of soft tick infestation. Condor 110:134-142.


Research on Atlantic Seabirds
World Seabird Union