Richard Boyles

Assistant Professor of Plant Breeding & Genetics
Plant and Environmental Sciences Department, PeeDee Research and Education Center

Office: 210 Pitner Center
Phone: 843-519-0488


 Educational Background

Ph.D. Genetics
Clemson University 2016

B.S. Biological Sciences
University of South Carolina 2012

 Research Interests

My research focuses on cereal crop improvement using a combination of traditional and advanced plant genetics and breeding approaches. My interests are (1) to develop crop varieties and hybrids with superior performance in the region, (2) to broaden the current gene pool that exists in regional breeding populations, and (3) to identify useful genes and alleles for important traits in grass row crops including grain yield and quality components.


Mural R, Grzybowski M, Mao C, Damke A, Sapkota S, Boyles R*, Salas-Fernandez M, Schnable P, Sigmon B, Kresovich S, Schnable J. Meta-analysis identifies pleiotropic loci controlling phenotypic trade-offs in sorghum. bioRxiv. doi: 10.1101/2020.10.27.355495.

Ibrahim A, Sutton R, Johnson J, Mergoum M, Simoneaux B, Harrison S, Murphy J, Mason R, Babar A, Neely C, Opena G, Jin Y, Kolmer J, Boyles R*, Cambron S, Baik B-K, Brown-Guedira G, Marshall D, Fountain M (2020) Registration of ‘GA06343-13E2 (TX-EL2)’ soft red winter wheat. Journal of Plant Registrations 15:107-112. doi: 10.1002/plr2.20031.

Disharoon A, Boyles R*, Jordan K, Kresovich S (2020) Exploring diverse sorghum (Sorghum bicolor (L.) Moench) accessions for malt amylase activity. Journal of the Institute of Brewing 127:5-12. doi: 10.1002/jib.628

Sapkota S, Boatwright J, Jordan K, Boyles R*, Kresovich S (2020) Identification of novel genomic associations and gene candidates for grain starch content in sorghum. Genes 11:1448. doi: 10.3390/genes11121448

Shields L, Gang Y, Sapkota S, Jordan K, Boatwright J, Jiang X, Kresovich S, Boyles R* (2020) Antimicrobial activity against foodborne pathogens in global sorghum [Sorghum bicolor (L.) Moench] germplasm. Crop Science. doi: 10.1002/csc2.20348

Sapkota S, Boatwright J, Jordan K, Boyles R*, Kresovich S (2020) Multi-trait regressor stacking increased genomic prediction accuracy of sorghum grain composition. Agronomy 10:1221. doi: 10.3390/agronomy10091221

Brenton Z, Juengst B, Cooper E, Myer M, Jordan K, Dale S, Glaubitz J, Wang X, Boyles R*, Connelly E, Kresovich S (2020) Species-specific duplication event confers adaptive benefits in Sorghum bicolor. G3: Genes|Genomes|Genetics. doi: 10.1534/g3.119.400921.

Sapkota S, Boyles R*, Cooper E, Brenton Z, Myers M, Kresovich S (2020) Impact of sorghum racial structure and diversity on genomic prediction of grain yield components. Crop Science. doi: 10.2135/cropsci2019.04.0221.

Boyles R*, Marshall D, Bockelman H (2019) Yield data from the Uniform Southern Soft Red Winter Wheat Nursery emphasize importance of selection location and environment for cultivar development. Crop Science 59: 1887-1898. doi: 10.2135/cropsci2018.11.0685.

Boyles R*, Brenton Z, Kresovich S (2019) Genetic and genomic resources of sorghum to connect genotype with phenotype in contrasting environments. The Plant Journal 97:19-39. doi: 10.1111/tpj.14113.

Boyles R*, Pfieffer B, Cooper E, Zielinski K, Myers M, Rooney W, and Kresovich S (2017) QTL mapping of agronomic and yield traits in two grain sorghum biparental families. Crop Science 57:2443-2456. doi: 10.2135/cropsci2016.12.0988.

Rhodes D, Hoffman Jr L, Rooney W, Herald T, Bean S, Brenton Z, Boyles R*, Kresovich S (2017) Genetic architecture of kernel composition in global sorghum germplasm. BMC Genomics 18:15 doi: 10.1186/s12864-016-3403-x.

Boyles R*, Pfieffer B, Rauh B, Cooper E, Zielinski K, Myers M, Brenton Z, Rooney W, Kresovich S (2016) Genetic dissection of sorghum grain quality traits using diverse and segregating populations. Theor Appl Genet doi: 10.1007/s00122-016-2844-6.

Boyles R*, Cooper E, Myers M, Brenton Z, Rauh B, Morris G, Kresovich S (2016) Genome-wide association studies of grain yield components in diverse sorghum germplasm. Plant Genome 9. doi: 10.3835/plantgenome2015.09.0091.

Brenton Z, Cooper E, Myers M, Boyles R*, Shakoor N, Zielinski K, Rauh B, Bridges W, Morris G, Kresovich S (2016) A genomic resource for the development, improvement, and exploitation of sorghum for bioenergy. Genetics genetics.115.183947. doi:10.1534/genetics.115.183947.

Shakoor N, Ziegler G, Dilkes B, Brenton Z, Boyles R*, Connolly E, Kresovich S, Baxter I (2016) Integration of experiments across diverse environments identifies the genetic determinants of variation in Sorghum bicolor seed element composition. Plant Physiol 170:1989–1998. doi:10.1104/pp.15.01971.

Kresovich S, Brenton Z, Myers M, Boyles R* (2014) Notice of release of nine cellulosic biomass sorghum hybrids: CU-C003, CU-C041, CU-C053, CU-C056, CU-C057, CU-C060, CU-C087, CU-C126, and CU-C225. Advanced Plant Technology Program, Clemson University.

Boyles R (2019) Exploring diversity and genomics-based approaches for sorghum yield gains. Center for Sorghum Improvement, Manhattan, KS.

Ackerman A, Boyles R* (2019) The Reestablished Clemson Small Grains Breeding Program. South Carolina Seedsmen Association annual meeting, Hilton Head, SC.

Boyles R (2019) Accelerating sorghum improvement with technological and genomics advances. Sorghum Production and Marketing Meeting, Columbia, SC.

Boyles R (2018) Improving agriculture with the power of genetics and breeding. SC Governor's School for Science and Mathematics, Hartsville, SC.

Boyles R*, Kresovich S (2018) Multi-population mapping data and their use in sorghum improvement. USDA NIFA FACT Data Driven Ag Workshop, Arlington, VA.

Boyles R (2018) Current status and future prospective of the Clemson University sorghum and small grains breeding program. SC Crop Improvement Association annual board meeting, Columbia, SC.

Kresovich S, Cooper E, Boyles R* (2017) The value of rediscovering heirlooms for the future of crop improvement and agriculture. HudsonAlpha CROPS Conference, Huntsville, AL.

Boyles R*, Cooper E, Brenton Z, Zielinski K, Rauh B, Morris G, Kresovich S (2016) Utilizing established sorghum mapping populations to identify QTL for grain yield and compositional traits. Plant and Animal Genome Annual Conference, San Diego, CA.

Boyles R*, Rhodes D, Shakoor N, Kresovich S (2014) Sorghum: The gluten-free, non-GMO, climate resilient cereal for the 21st century. ASA, CSSA, & SSSA International Annual Meeting, Long Beach, CA.

Boyles R*, Rooney W, Myers M, Brenton Z, Kresovich S (2014) The Genetic and Biochemical Frameworks Influencing the Energy Value of Sorghum Grain. ASA, CSSA, & SSSA International Annual Meeting, Long Beach, CA.

Boyles R*, Kresovich S (2014) Advanced Plant Technology Program. South Carolina Seedsmen Association annual meeting, Hilton Head, SC.

Boyles R*, Morris G, Rhodes D, Hoffman Jr L, Rooney W, Kresovich S (2013) Genome-wide Association Studies of Grain Traits in Sorghum [Sorghum bicolor (L.) Moench]. ASA, CSSA, & SSSA International Annual Meeting, Tampa, FL.


OREI Project Website
Wheat historical selection impacts
Sorghum genetic and genomic resources
Sorghum grain quality dissection
GWAS of grain yield components